Vishwa T. Kasoju

Bio: Vishwa T. Kasoju is an academic researcher from Oklahoma State University–Stillwater. The author has contributed to research in topics: Wing & Drag. The author has an hindex of 4, co-authored 10 publications receiving 63 citations.

Leaky Flow through Simplified Physical Models of Bristled Wings of Tiny Insects during Clap and Fling

Abstract: In contrast to larger flight-capable insects such as hawk moths and fruit flies, miniature flying insects such as thrips show the obligatory use of wing–wing interaction via “clap and fling” during the end of upstroke and start of downstroke. Although fling can augment lift generated during flapping flight at chord-based Reynolds number (Re) of 10 or lower, large drag forces are necessary to clap and fling the wings. In this context, bristles observed in the wings of most tiny insects have been shown to lower drag force generated in clap and fling. However, the fluid dynamic mechanism underlying drag reduction by bristled wings and the impact of bristles on lift generated via clap and fling remain unclear. We used a dynamically scaled robotic model to examine the forces and flow structures generated during clap and fling of: three bristled wing pairs with varying inter-bristle spacing, and a geometrically equivalent solid wing pair. In contrast to the solid wing pair, reverse flow through the gaps between the bristles was observed throughout clap and fling, resulting in: (a) drag reduction; and (b) weaker and diffuse leading edge vortices that lowered lift. Shear layers were formed around the bristles when interacting bristled wing pairs underwent clap and fling motion. These shear layers lowered leakiness of flow through the bristles and minimized loss of lift in bristled wings. Compared to the solid wing, peak drag coefficients were reduced by 50–90% in bristled wings. In contrast, peak lift coefficients of bristled wings were only reduced by 35–60% from those of the solid wing. Our results suggest that the bristled wings can provide unique aerodynamic benefits via increasing lift to drag ratio during clap and fling for Re between 5 and 15.

Flow Structure and Force Generation on Flapping Wings at Low Reynolds Numbers Relevant to the Flight of Tiny Insects

Abstract: In contrast to larger species, little is known about the flight of the smallest flying insects, such as thrips and fairyflies. These tiny animals range from 300 to 1000 microns in length and fly at Reynolds numbers ranging from about 4 to 60. Previous work with numerical and physical models have shown that the aerodynamics of these diminutive insects is significantly different from that of larger animals, but most of these studies have relied on two-dimensional approximations. There can, however, be significant differences between two- and three-dimensional flows, as has been found for larger insects. To better understand the flight of the smallest insects, we have performed a systematic study of the forces and flow structures around a three-dimensional revolving elliptical wing. We used both a dynamically scaled physical model and a three-dimensional computational model at Reynolds numbers ranging from 1 to 130 and angles of attacks ranging from 0° to 90°. The results of the physical and computational models were in good agreement and showed that dimensionless drag, aerodynamic efficiency, and spanwise flow all decrease with decreasing Reynolds number. In addition, both the leading and trailing edge vortices remain attached to the wing over the scales relevant to the smallest flying insects. Overall, these observations suggest that there are drastic differences in the aerodynamics of flight at the scale of the smallest flying animals.

Aerodynamic effects of varying solid surface area of bristled wings performing clap and fling.

Abstract: The smallest flying insects with body lengths under 2 mm show a marked preference for wings consisting of a thin membrane with long bristles, and the use of clap and fling kinematics to augment lift at Reynolds numbers (Re) of approximately 10. Bristled wings have been shown to reduce drag forces in clap and fling, but the aerodynamic roles of several bristled wing geometric variables remain unclear. This study examines the effects of varying the ratio of membrane area (A M) to total wing area (A T) on aerodynamic forces and flow structures generated during clap and fling at Re on the order of 10. We also examine the aerodynamic consequences of scaling bristled wings to Re = 120, relevant to flight of fruit flies. We analyzed published forewing images of 25 species of thrips (Thysanoptera) and found that A M/A T ranged from 14% to 27%, as compared to 11% to 88% previously reported for smaller-sized fairyflies (Hymenoptera). These data were used to develop physical bristled wing models with A M/A T ranging from 15% to 100%, which were tested in a dynamically scaled robotic clap and fling model. At all Re, bristled wings produced slightly lower lift coefficients (C L) when compared to solid wings, but provided significant drag reduction. At Re = 10, largest values of peak lift over peak drag ratios were generated by wing models with A M/A T similar to thrips forewings (15% to 30%). Circulation of the leading edge vortex and trailing edge vortex decreased with decreasing A M/A T during clap and fling at Re = 10. Decreased chordwise circulation near the wing tip, vortex shedding, and interaction between flow structures from clap with those from fling resulted in lowering C L generated via clap and fling at Re = 120 as compared to Re = 10. Clap and fling becomes less beneficial at Re = 120, regardless of the drag reduction provided by bristled wings.

Aerodynamic interaction of bristled wing pairs in fling

Abstract: Tiny flying insects of body lengths under 2 mm use the “clap-and-fling” mechanism with bristled wings for lift augmentation and drag reduction at a chord-based Reynolds number (Re) on O ( 10 ). We examine the wing–wing interaction of bristled wings in fling at Re = 10 as a function of initial inter-wing spacing (δ) and degree of overlap between rotation and linear translation. A dynamically scaled robotic platform was used to drive physical models of bristled wing pairs with the following kinematics (all angles relative to vertical): (1) rotation about the trailing edge to angle θr, (2) linear translation at a fixed angle (θt), and (3) combined rotation and linear translation. The results show that (1) the cycle-averaged drag coefficient decreased with increasing θr and θt and (2) decreasing δ increased the lift coefficient owing to increased asymmetry in the circulation of leading and trailing edge vortices. A new dimensionless index, reverse flow capacity (RFC), was used to quantify the maximum possible ability of a bristled wing to leak the fluid through the bristles. The drag coefficients were larger for smaller δ and θr despite larger RFC, likely due to the blockage of inter-bristle flow by shear layers around the bristles. Smaller δ during early rotation resulted in the formation of strong positive pressure distribution between the wings, resulting in an increased drag force. The positive pressure region weakened with increasing θr, which in turn reduced the drag force. Tiny insects have been previously reported to use large rotational angles in fling, and our findings suggest that a plausible reason is to reduce drag forces.

Aerodynamic interaction of bristled wing pairs in fling

Abstract: Tiny flying insects of body lengths under 2 mm use the `clap-and-fling' mechanism with bristled wings for lift augmentation and drag reduction at chord-based Reynolds number ($Re$) on $\mathcal{O}$(10). We examine wing-wing interaction of bristled wings in fling at $Re$=10, as a function of initial inter-wing spacing ($\delta$) and degree of overlap between rotation and linear translation. A dynamically scaled robotic platform was used to drive physical models of bristled wing pairs with the following kinematics (all angles relative to vertical): 1) rotation about the trailing edge to angle $\theta_\text{r}$; 2) linear translation at a fixed angle ($\theta_\text{t}$); and 3) combined rotation and linear translation. The results show that: 1) cycle-averaged drag coefficient decreased with increasing $\theta_\text{r}$ and $\theta_\text{t}$; and 2) decreasing $\delta$ increased the lift coefficient owing to increased asymmetry in circulation of leading and trailing edge vortices. A new dimensionless index, reverse flow capacity (RFC), was used to quantify the maximum possible ability of a bristled wing to leak fluid through the bristles. Drag coefficients were larger for smaller $\delta$ and $\theta_\text{r}$ despite larger RFC, likely due to blockage of inter-bristle flow by shear layers around the bristles. Smaller $\delta$ during early rotation resulted in formation of strong positive pressure distribution between the wings, resulting in increased drag force. The positive pressure region weakened with increasing $\theta_\text{r}$, which in turn reduced drag force. Tiny insects have been reported to use large rotational angles in fling, and our findings suggest that a plausible reason is to reduce drag forces.

Mimicking insect wings: The roadmap to bioinspiration

Abstract: Insect wings possess unique, multifaceted properties that have drawn increasing attention in recent times. They serve as an inspiration for engineering of materials with exquisite properties. The structure–function relationships of insect wings are yet to be documented in detail. In this review, we present a detailed understanding of the multifunctional properties of insect wings, including micro- and nanoscale architecture, material properties, aerodynamics, sensory perception, wettability, optics, and antibacterial activity, as investigated by biologists, physicists, and engineers. Several established modeling strategies and fabrication methods are reviewed to engender novel ideas for biomimetics in diverse areas.

Novel flight style and light wings boost flight performance of tiny beetles

Abstract: Flight speed is positively correlated with body size in animals1. However, miniature featherwing beetles can fly at speeds and accelerations of insects three times their size2. Here we show that this performance results from a reduced wing mass and a previously unknown type of wing-motion cycle. Our experiment combines three-dimensional reconstructions of morphology and kinematics in one of the smallest insects, the beetle Paratuposa placentis (body length 395 μm). The flapping bristled wings follow a pronounced figure-of-eight loop that consists of subperpendicular up and down strokes followed by claps at stroke reversals above and below the body. The elytra act as inertial brakes that prevent excessive body oscillation. Computational analyses suggest functional decomposition of the wingbeat cycle into two power half strokes, which produce a large upward force, and two down-dragging recovery half strokes. In contrast to heavier membranous wings, the motion of bristled wings of the same size requires little inertial power. Muscle mechanical power requirements thus remain positive throughout the wingbeat cycle, making elastic energy storage obsolete. These adaptations help to explain how extremely small insects have preserved good aerial performance during miniaturization, one of the factors of their evolutionary success.

Hydrodynamics of metachronal paddling: effects of varying Reynolds number and phase lag.

Abstract: Negatively buoyant freely swimming crustaceans such as krill must generate downward momentum in order to maintain their position in the water column. These animals use a drag-based propulsion strategy, where pairs of closely spaced swimming limbs are oscillated rhythmically from the tail to head. Each pair is oscillated with a phase delay relative to the neighbouring pair, resulting in a metachronal wave travelling in the direction of animal motion. It remains unclear how oscillations of limbs in the horizontal plane can generate vertical momentum. Using particle image velocimetry measurements on a robotic model, we observed that metachronal paddling with non-zero phase lag created geometries of adjacent paddles that promote the formation of counter-rotating vortices. The interaction of these vortices resulted in generating large-scale angled downward jets. Increasing phase lag resulted in more vertical orientation of the jet, and phase lags in the range used by Antarctic krill produced the most total momentum. Synchronous paddling produced lower total momentum when compared with metachronal paddling. Lowering Reynolds number by an order of magnitude below the range of adult krill (250-1000) showed diminished downward propagation of the jet and lower vertical momentum. Our findings show that metachronal paddling is capable of producing flows that can generate both lift (vertical) and thrust (horizontal) forces needed for fast forward swimming and hovering.

Aerodynamic performance of a bristled wing of a very small insect

Abstract: Aerodynamic force generation capacity of the wing of a miniature beetle Paratuposa placentis is evaluated using a combined experimental and numerical approach. The wing has a peculiar shape reminiscent of a bird feather, often found in the smallest insects. Aerodynamic force coefficients are determined from a dynamically scaled force measurement experiment with rotating bristled and membrane wing models in a glycerin tank. Subsequently, they are used as numerical validation data for computational fluid dynamics simulations using an adaptive Navier–Stokes solver. The latter provides access to important flow properties such as leakiness and permeability. It is found that, in the considered biologically relevant regimes, the bristled wing functions as a less than $$50\%$$ leaky paddle, and it produces between 66 and $$96\%$$ of the aerodynamic drag force of an equivalent membrane wing. The discrepancy increases with increasing Reynolds number. It is shown that about half of the aerodynamic normal force exerted on a bristled wing is due to viscous shear stress. The paddling effectiveness factor is proposed as a measure of aerodynamic efficiency.