Showing papers by "San Francisco State University published in 2002"

Network structure and biodiversity loss in food webs: robustness increases with connectance

Abstract: Food-web structure mediates dramatic effects of biodiversity loss including secondary and ‘cascading’ extinctions. We studied these effects by simulating primary species loss in 16 food webs from terrestrial and aquatic ecosystems and measuring robustness in terms of the secondary extinctions that followed. As observed in other networks, food webs are more robust to random removal of species than to selective removal of species with the most trophic links to other species. More surprisingly, robustness increases with food-web connectance but appears independent of species richness and omnivory. In particular, food webs experience ‘rivet-like’ thresholds past which they display extreme sensitivity to removal of highly connected species. Higher connectance delays the onset of this threshold. Removing species with few trophic connections generally has little effect though there are several striking exceptions. These findings emphasize how the number of species removed affects ecosystems differently depending on the trophic functions of species removed.

Food-web structure and network theory: The role of connectance and size.

Abstract: Networks from a wide range of physical, biological, and social systems have been recently described as “small-world” and “scale-free.” However, studies disagree whether ecological networks called food webs possess the characteristic path lengths, clustering coefficients, and degree distributions required for membership in these classes of networks. Our analysis suggests that the disagreements are based on selective use of relatively few food webs, as well as analytical decisions that obscure important variability in the data. We analyze a broad range of 16 high-quality food webs, with 25–172 nodes, from a variety of aquatic and terrestrial ecosystems. Food webs generally have much higher complexity, measured as connectance (the fraction of all possible links that are realized in a network), and much smaller size than other networks studied, which have important implications for network topology. Our results resolve prior conflicts by demonstrating that although some food webs have small-world and scale-free structure, most do not if they exceed a relatively low level of connectance. Although food-web degree distributions do not display a universal functional form, observed distributions are systematically related to network connectance and size. Also, although food webs often lack small-world structure because of low clustering, we identify a continuum of real-world networks including food webs whose ratios of observed to random clustering coefficients increase as a power–law function of network size over 7 orders of magnitude. Although food webs are generally not small-world, scale-free networks, food-web topology is consistent with patterns found within those classes of networks.

Genome sequence of the plant pathogen Ralstonia solanacearum

Abstract: Ralstonia solanacearum is a devastating, soil-borne plant pathogen with a global distribution and an unusually wide host range. It is a model system for the dissection of molecular determinants governing pathogenicity. We present here the complete genome sequence and its analysis of strain GMI1000. The 5.8-megabase (Mb) genome is organized into two replicons: a 3.7-Mb chromosome and a 2.1-Mb megaplasmid. Both replicons have a mosaic structure providing evidence for the acquisition of genes through horizontal gene transfer. Regions containing genetically mobile elements associated with the percentage of G+C bias may have an important function in genome evolution. The genome encodes many proteins potentially associated with a role in pathogenicity. In particular, many putative attachment factors were identified. The complete repertoire of type III secreted effector proteins can be studied. Over 40 candidates were identified. Comparison with other genomes suggests that bacterial plant pathogens and animal pathogens harbour distinct arrays of specialized type III-dependent effectors.

Closing the seed dispersal loop

Abstract: Seed dispersal links the end of the reproductive cycle of adult plants with the establishment of their offspring, and is widely accepted to have a profound effect on vegetation structure. Confirming and quantifying this effect, however, has proven to be a challenge. Recent research on animal-mediated seed dispersal has brought us closer to this goal: ecologists have been explicitly examining the processes that mediate seed deposition and seedling recruitment. Exciting new techniques, such as the analysis of stable isotope ratios and molecular genetic markers, are making it possible to relate dispersed seeds and seedlings back to parent plants. Meanwhile, evidence from plant demography research is revealing that seed dispersal might have an important role in determining patterns of tree diversity and distribution. The continued synergy between seed dispersal research and the study of plant demography should help researchers link seed dispersal and adult vegetation structure, closing the seed dispersal loop.

Dinitrogen fixation in the world's oceans

Abstract: The surface water of the marine environment has traditionally been viewed as a nitrogen (N) limited habitat, and this has guided the development of conceptual biogeochemical models focusing largely on the reservoir of nitrate as the critical source of N to sustain primary productivity. However, selected groups of Bacteria, including cyanobacteria, and Archaea can utilize dinitrogen (N2) as an alternative N source. In the marine environment, these microorganisms can have profound effects on net community production processes and can impact the coupling of C-N-P cycles as well as the net oceanic sequestration of atmospheric carbon dioxide. As one component of an integrated ‘Nitrogen Transport and Transformations’ project, we have begun to re-assess our understanding of (1) the biotic sources and rates of N2 fixation in the world’s oceans, (2) the major controls on rates of oceanic N2 fixation, (3) the significance of this N2 fixation for the global carbon cycle and (4) the role of human activities in the alteration of oceanic N2 fixation. Preliminary results indicate that rates of N2 fixation, especially in subtropical and tropical open ocean habitats, have a major role in the global marine N budget. Iron (Fe) bioavailability appears to be an important control and is, therefore, critical in extrapolation to global rates of N2 fixation. Anthropogenic perturbations may alter N2 fixation in coastal environments through habitat destruction and eutrophication, and open ocean N2 fixation may be enhanced by warming and increased stratification of the upper water column. Global anthropogenic and climatic changes may also affect N2 fixation rates, for example by altering dust inputs (i.e. Fe) or by expansion of subtropical boundaries. Some recent estimates of global ocean N2 fixation are in the range of 100−200 Tg N (1−2 × 1014 g N) yr −1, but have large uncertainties. These estimates are nearly an order of magnitude greater than historical, pre-1980 estimates, but approach modern estimates of oceanic denitrification.

Arsenic(III) Oxidation and Arsenic(V) Adsorption Reactions on Synthetic Birnessite

Abstract: The oxidation of arsenite (As(III)) by manganese oxide is an important reaction in both the natural cycling of As and the development of remediation technology for lowering the concentration of dissolved As(III) in drinking water. This study used both a conventional stirred reaction apparatus and extended X-ray absorption fine structure (EXAFS) spectroscopy to investigate the reactions of As(III) and As(V) with synthetic birnessite (MnO2). Stirred reactor experiments indicate that As(III) is oxidized by MnO2 followed by the adsorption of the As(V) reaction product on the MnO2 solid phase. The As(V)-Mn interatomic distance determined by EXAFS analysis for both As(III)- and As(V)-treated MnO2 was 3.22 A, giving evidence for the formation of As(V) adsorption complexes on MnO2 crystallite surfaces. The most likely As(V)-MnO2 complex is a bidentate binuclear corner sharing (bridged) complex occurring at MnO2 crystallite edges and interlayer domains. In the As(III)-treated MnO2 systems, reductive dissolution of the MnO2 solid during the oxidation of As(III) caused an increase in the adsorption of As(V) when compared with As(V)-treated MnO2. This suggested that As(III) oxidation caused a surface alteration, creating fresh reaction sites for As(V) on MnO2 surfaces.

Radial Velocities for 889 Late-Type Stars*

Abstract: We report radial velocities for 844 FGKM-type main-sequence and subgiant stars and 45 K giants, most of which had either low-precision velocity measurements or none at all. These velocities differ from the stand- ard stars of Udry et al. by 0.035 km s � 1 (rms) for the 26 FGK standard stars in common. The zero point of our velocities differs from that of Udry et al.: hVPresentVUdry i¼þ 0:053 km s � 1 . Thus, these new velocities agree with the best known standard stars both in precision and zero point, to well within 0.1 km s � 1 . Nonetheless, both these velocities and the standards suffer from three sources of systematic error, namely, convective blueshift, gravitational redshift, and spectral type mismatch of the reference spectrum. These sys- tematic errors are here forced to be zero for G2 V stars by using the Sun as reference, with Vesta and day sky as proxies. But for spectral types departing from solar, the systematic errors reach 0.3 km s � 1 in the F and K stars and 0.4 km s � 1 in M dwarfs. Multiple spectra were obtained for all 889 stars during 4 years, and 782 of them exhibit velocity scatter less than 0.1 km s � 1 . These stars may serve as radial velocity standards if they remain constant in velocity. We found 11 new spectroscopic binaries and report orbital parameters for them. Subject headings: binaries: spectroscopic — catalogs — stars: fundamental parameters — stars: kinematics — stars: late-type — techniques: radial velocities — techniques: spectroscopic On-line material: machine-readable tables

Bird song, ecology and speciation

Abstract: The study of bird song dialects was once considered the most promising approach for investigating the role of behaviour in reproductive divergence and speciation. However, after a series of studies yielding conflicting results, research in the field slowed significantly. Recent findings, on how ecological factors may lead to divergence in both song and morphology, necessitate a re-examination. We focus primarily on species with learned song, examine conflicting results in the literature and propose some potential new directions for future studies. We believe an integrative approach, including an examination of the role of ecology in divergent selection, is essential for gaining insight into the role of song in the evolution of assortative mating. Habitat-dependent selection on both song and fitness-related characteristics can lead to parallel divergence in these traits. Song may, therefore, provide females with acoustic cues to find males that are most fit for a particular habitat. In analysing the role of song learning in reproductive divergence, we focus on post-dispersal plasticity in a conceptual framework. We argue that song learning may initially constrain reproductive divergence, while in the later stages of population divergence it may promote speciation.

Applying Covariational Reasoning While Modeling Dynamic Events: A Framework and a Study

Abstract: The article develops the notion of covariational reasoning and proposes a framework for describing the mental actions involved in applying covariational reasoning when interpreting and representing dynamic function events. It also reports on an investigation of high-performing 2nd-semester calculus students’ ability to reason about covarying quantities in dynamic situations. The study revealed that these students were able to construct images of a function’s dependent variable changing in tandem with the imagined change of the independent variable, and in some situations, were able to construct images of rate of change for contiguous intervals of a function’s domain. However, students appeared to have difficulty forming images of continuously changing rate and could not accurately represent or interpret inflection points or increasing and decreasing rate for dynamic function situations. These findings suggest that curriculum and instruction should place increased emphasis on moving students from a coordinated image of two variables changing in tandem to a coordinated image of the instantaneous rate of change with continuous changes in the independent variable for dynamic function situations.

Arsenic(III) and arsenic(V) reactions with zerovalent iron corrosion products.

Abstract: Zerovalent iron (Fe0) has tremendous potential as a remediation material for removal of arsenic from groundwater and drinking water. This study investigates the speciation of arsenate (As(V)) and arsenite (As(III)) after reaction with two Fe0 materials, their iron oxide corrosion products, and several model iron oxides. A variety of analytical techniques were used to study the reaction products including HPLC-hydride generation atomic absorption spectrometry, X-ray diffraction, scanning electron microscopy-energy-dispersive X-ray analysis, and X-ray absorption spectroscopy. The products of corrosion of Fe0 include lepidocrocite (γ-FeOOH), magnetite (Fe3O4), and/or maghemite (γ-Fe2O3), all of which indicate Fe(II) oxidation as an intermediate step in the Fe0 corrosion process. The in-situ Fe0 corrosion reaction caused a high As(III) and As(V) uptake with both Fe0 materials studied. Under aerobic conditions, the Fe0 corrosion reaction did not cause As(V) reduction to As(III) but did cause As(III) oxidation ...

Two degrees of separation in complex food webs.

Abstract: Feeding relationships can cause invasions, extirpations, and population fluctuations of a species to dramatically affect other species within a variety of natural habitats. Empirical evidence suggests that such strong effects rarely propagate through food webs more than three links away from the initial perturbation. However, the size of these spheres of potential influence within complex communities is generally unknown. Here, we show for that species within large communities from a variety of aquatic and terrestrial ecosystems are on average two links apart, with >95% of species typically within three links of each other. Species are drawn even closer as network complexity and, more unexpectedly, species richness increase. Our findings are based on seven of the largest and most complex food webs available as well as a food-web model that extends the generality of the empirical results. These results indicate that the dynamics of species within ecosystems may be more highly interconnected and that biodiversity loss and species invasions may affect more species than previously thought.

Nitrogen Fixation and Nitrogen Isotope Abundances in Zooplankton of the Oligotrophic North Atlantic

Abstract: Deep-water nitrate is a major reservoir of oceanic combined nitrogen and has long been considered to be the major source of new nitrogen supporting primary production in the oligotrophic ocean. 15N:14N ratios in plankton provide an integrative record of the nitrogen cycle processes at work in the ocean, and near-surface organic matter in oligotrophic waters like the Sargasso Sea is characterized by an unusually low 15N content relative to average deep-water nitrate. Herein we show that the low dΔ15N of suspended particles and zooplankton from the tropical North Atlantic cannot arise through isotopic fractionation associated with nutrient uptake and food web processes but are instead consistent with a significant input of new nitrogen to the upper water column by N2 fixation. These results provide direct, integrative evidence that N2 fixation makes a major contribution to the nitrogen budget of the oligotrophic North Atlantic Ocean.

Effects of freshwater flow on abundance of estuarine organisms : physical effects or trophic linkages?

Abstract: All ecosystems are influenced by physical forcing. Estuarine ecosystems respond most strongly on an interannual timescale to variability in freshwater flow. Several mechanisms for positive or negative flow effects on biological populations in estuaries have been proposed; however, positive effects appear to operate mainly through stimulation of primary production with effects propagating up the food web. In the northern San Francisco Estuary, abundance or survival of several common species of fish and shrimp varied positively with flow-in data through 1992. I re-examined these relationships and those of several additional taxa in an analysis of long-term (20 to 40 yr) mon- itoring data. The spread of the introduced clam Potamocorbula amurensis in 1987 provided an oppor- tunity to examine simultaneously the responses of estuarine species to flow and to changes in the food web. I separated variability into a flow response, a step change after 1987 and other sources of variability. Responses of fish and shrimp contrasted with those of lower trophic levels. All but 1 spe- cies of nekton responded positively to flow, only 2 had clear declines after 1987, and none of the rela- tionships changed in slope after 1987. In contrast with the higher trophic levels, chlorophyll a (chl a) and several species of zooplankton declined markedly after 1987, and had either weak responses to flow or responses that changed after 1987. Thus, the food web appears strongly coupled between benthos and plankton, and weakly coupled between zooplankton and fish, as has been found in other systems. More importantly, the variation with freshwater flow of abundance or survival of organisms in higher trophic levels apparently did not occur through upward trophic transfer, since a similar rela- tionship was lacking in most of the data on lower trophic levels. Rather, this variation may occur through attributes of physical habitat that vary with flow.

Habitat-dependent song divergence in the little greenbul: an analysis of environmental selection pressures on acoustic signals

Abstract: Bird song is a sexual trait important in mate choice and known to be shaped by environmental selection. Here we investigate the ecological factors shaping song variation across a rainforest gradient in central Africa. We show that the little greenbul (Andropadus virens), previously shown to vary morphologically across the gradient in fitness-related characters, also varies with respect to song characteristics. Acoustic features, including minimum and maximum frequency, and delivery rate of song notes showed significant differences between habitats. In contrast, we found dialectal variation independent of habitat in population-typical songtype sequences. This pattern is consistent with ongoing gene flow across habitats and in line with the view that song variation in the order in which songtypes are produced is not dependent on habitat characteristics in the same way physical song characteristics are. Sound transmission characteristics of the two habitats did not vary significantly, but analyses of ambient noise spectra revealed dramatic and consistent habitat-dependent differences. Matching between low ambient noise levels for low frequencies in the rainforest and lower minimal frequencies in greenbul songs in this habitat suggests that part of the song divergence may be driven by habitat-dependent ambient noise patterns. These results suggest that habitat-dependent selection may act simultaneously on traits of ecological importance and those important in prezygotic isolation, leading to an association between morphological and acoustic divergence. Such an association may promote assortative mating and may be a mechanism driving reproductive divergence across ecological gradients.

Environmental Justice and Regional Inequality in Southern California: Implications for Future Research

Abstract: Environmental justice offers researchers new insights into the juncture of social inequality and public health and provides a framework for policy discussions on the impact of discrimination on the environmental health of diverse communities in the United States. Yet, causally linking the presence of potentially hazardous facilities or environmental pollution with adverse health effects is difficult, particularly in situations in which diverse populations are exposed to complex chemical mixtures. A community-academic research collaborative in southern California sought to address some of these methodological challenges by conducting environmental justice research that makes use of recent advances in air emissions inventories and air exposure modeling data. Results from several of our studies indicate that communities of color bear a disproportionate burden in the location of treatment, storage, and disposal facilities and Toxic Release Inventory facilities. Longitudinal analysis further suggests that facility siting in communities of color, not market-based "minority move-in," accounts for these disparities. The collaborative also investigated the health risk implications of outdoor air toxics exposures from mobile and stationary sources and found that race plays an explanatory role in predicting cancer risk distributions among populations in the region, even after controlling for other socioeconomic and demographic indicators. Although it is unclear whether study results from southern California can be meaningfully generalized to other regions in the United States, they do have implications for approaching future research in the realm of environmental justice. The authors propose a political economy and social inequality framework to guide future research that could better elucidate the origins of environmental inequality and reasons for its persistence.

One-dimensional ecosystem model of the equatorial Pacific upwelling system. Part I: model development and silicon and nitrogen cycle

Abstract: A one-dimensional ecosystem model was developed for the equatorial Pacific upwelling system, and the model was used to study nitrogen and silicon cycle in the equatorial Pacific. The ecosystem model consisted of 10 components (nitrate, silicate, ammonium, small phytoplankton, diatom, micro- and meso-zooplankton, detrital nitrogen and silicon, and total CO2). The ecosystem model was forced by the area-averaged (5°S–5°N, 90°W–180°, the Wyrtki Box) annual mean upwelling velocity and vertical diffusivity obtained from a three-dimensional circulation model. The model was capable of reproducing the low-silicate, high-nitrate, and low-chlorophyll (LSHNLC) conditions in the equatorial Pacific. The linkage to carbon cycle was through the consumption of assimilated nitrate and silicate (i.e. new productions). Model simulations demonstrated that low-silicate concentration in the equatorial Pacific limits production of diatoms, and it resulted in low percentage of diatoms, 16%, in the total phytoplankton biomass. In the area of 5°S–5°N and 90°W–180°, the model produced an estimated sea-to-air CO2 flux of 4.3 mol m−2 yr−1, which is consistent with the observed results ranging of 1.0–4.5 mol m−2 yr−1. The ammonium inhibition played an important role in determining the nitrogen cycle in the model. The modeled surface nitrate concentration could increase by a factor of 10 (from 0.8 to 8.0 mmol m−3) when the strength of the ammonium inhibition increased from ψ=1.0 to 10.0 (mmol m−3)–1. The effects of both micro- and meso-zooplankton grazing were tested by varying the micro- and meso-zooplankton maximum grazing rates, G1max and G2max. The modeled results were quite sensitive to the zooplankton grazing parameters. The current model considered the role of iron implicitly through the parameters that determine the growth rate of diatoms. Several iron-enrichment experiments were conducted by changing the parameter α (the initial slope of the photosynthetic rate over irradiance at low irradiance), KSi(OH)4 (half-saturation concentration of silicate uptake by diatom), and μ2max (the potential maximum specific diatom growth rate) in the regulation terms of silicate uptake by diatom. Within the first 5 days in the modeled iron-enrichment experiment, the diatom biomass increased from 0.08 to 2.5 mmol m−3, more than a factor of 30 increase. But the diatom populations crashed 2 weeks after the experiment started, due to exhaustion of available silicate and increased mesozooplankton population. The modeled iron-enrichment experiments produced several ecological behaviors similar to these observed during the IronEx-2.

Structure and Flow-Induced Variability of the Subtidal Salinity Field in Northern San Francisco Bay

Abstract: The structure of the salinity field in northern San Francisco Bay and how it is affected by freshwater flow are discussed. Two datasets are examined: the first is 23 years of daily salinity data taken by the U.S. Bureau of Reclamation along the axis of northern San Francisco Bay; the second is a set of salinity transects taken by the U.S. Geological Survey between 1988 and 1993. Central to this paper is a measure of salinity intrusion, X2: the distance from the Golden Gate Bridge to where the bottom salinity is 2 psu. Using X2 to scale distance, the authors find that for most flow conditions, the mean salinity distribution of the estuary is nearly self-similar with a salinity gradient in the center 70% of the region between the Golden Gate and X2 that is proportional to X−12. Analysis of covariability of Q and X2 showed a characteristic timescale of adjustment of the salinity field of approximately 2 weeks. The steady-state response deduced from the X2 time series implies that X2 is proportional ...

Dynamically close galaxy pairs and merger rate evolution in the cnoc2 redshift survey

Abstract: We investigate redshift evolution in the galaxy merger and accretion rates, using a well-defined sample of 4184 galaxies with 0.12 ≤ z ≤ 0.55 and RC ≤ 21.5. We identify 88 galaxies in close (5 ≤ rp ≤ 20 h-1 kpc) dynamical (Δv ≤ 500 km s-1) pairs. These galaxies are used to compute global pair statistics, after accounting for selection effects resulting from the flux limit, k-corrections, luminosity evolution, and spectroscopic incompleteness. We find that the number of companions per galaxy (for -21 ≤ M ≤ -18) is Nc = 0.0321 ± 0.0077 at z = 0.3. The luminosity in companions, per galaxy, is Lc = 0.0294 ± 0.0084 × 1010 h2 L☉. We assume that Nc is proportional to the galaxy merger rate, while Lc is directly related to the mass accretion rate. After increasing the maximum pair separation to 50 h-1 kpc and comparing with the low-redshift SSRS2 pair sample, we infer evolution in the galaxy merger and accretion rates of (1 + z)2.3±0.7 and (1 + z)2.3±0.9, respectively. These are the first such estimates to be made using only confirmed dynamical pairs. When combined with several additional assumptions, this implies that approximately 15% of present epoch galaxies with -21 ≤ MB ≤ -18 have undergone a major merger since z = 1.

Physical, Biological, and Management Responses to Variable Freshwater Flow into the San Francisco Estuary

Abstract: Freshwater flow is the principal cause of physical variability in estuaries and a focus of conflict in estuaries where a substantial fraction of the freshwater is diverted. Variation in freshwater flow can have many effects: inundation of flood plains, increase loading and advective transport of materials and organisms, dilution or mobilization of contam- inants, compression of the estuarine salinity field and density gradient, increase in stratification, and decrease in residence time for water while increasing it for some particles and biota. In the San Francisco Estuary, freshwater flow is highly variable, and has been altered by shifts in seasonal patterns of river flow and increases in diversions from tidal and non- tidal regions, entraining fish of several species of concern. Abundance or survival of several estuarine-dependent species also increases with freshwater outflow. These relationships to flow may be due to several potential mechanisms, each with its own locus and period of effectiveness, but no mechanism has been conclusively shown to underlie the flow relationship of any species. Several flow-based management actions were established in the mid-1990s, including a salinity standard based on these flow effects, as well as reductions in diversion pumping during critical periods for listed species of fish. The effectiveness of these actions has not been established. To make the salinity standard more effective and more applicable to future estuarine conditions will require investigation to determine the underlying mechanisms. Effects of entrainment at diversion facilities are more straightforward conceptually but difficult to quantify, and resolving these may require experimental manipulations of diversion flow.

Not as the crow flies: a historical explanation for circuitous migration in Swainson's thrush (Catharus ustulatus).

Abstract: Many migratory songbirds follow circuitous migratory routes instead of taking the shortest path between overwintering and breeding areas. Here, we study the migration patterns in Swainson's thrush (Catharus ustulatus), a neartic-neotropical migrant songbird, using molecular genetic approaches. This species is presently separated into genetically distinct coastal and continental populations that diverged during the Late Pleistocene (as indicated by molecular dating), yet appear to have retained ancestral patterns of migration. Low nucleotide diversity, a star-like haplotype phylogeny and unimodal mismatch distributions all support the hypothesis that both the coastal and the continental populations have undergone recent demographic expansions. Nearctic-neotropical banding and genetic data show nearly complete segregation of migratory routes and of overwintering locations: coastal populations migrate along the Pacific Coast to overwintering sites in Central America and Mexico, whereas continental populations migrate along an eastern route to overwintering sites in Panama and South America. Nearctic-neotropical banding data also show that continental birds north, northwest and east of this migratory divide fly thousands of miles east before turning south. We conclude that circuitous migration in the Swainson's thrush is an artefact of a Late Pleistocene range expansion.

The Relative Contribution of War Experiences and Exile-Related Stressors to Levels of Psychological Distress Among Bosnian Refugees

Abstract: This study examined the relative contribution of 2 exile-related variables—social isolation and daily activity level—and war experiences of violence and loss, to levels of PTSD and depressive symptomatology in 2 groups of Bosnian refugees, 1 clinical group (N = 59) and the other a nonclinical community (N = 40) group. As hypothesized, exposure to war-related violence was highly predictive of PTSD symptoms in both groups; in addition, social isolation was significantly related to PTSD symptomatology in the community group. In contrast, depressive symptomatology was accounted for primarily by the exile-related stressors. For the clinical group, depressive symptoms were also accounted for by experiences of war-related loss. The implications of these findings for mental health interventions with refugees are considered.

A Planet at 5 AU around 55 Cancri

Abstract: We report precise Doppler-shift measurements of 55 Cancri (G8 V) obtained from 1989 to 2002 at Lick Observatory. The velocities reveal evidence for an outer planetary companion to 55 Cancri orbiting at 5.5 AU. The velocities also conirm a second, inner planet at 0.11 AU. The outer planet is the irst extrasolar planet found that orbits near or beyond the orbit of Jupiter. It was drawn from a sample of 50 stars observed with suci ent duration and quality to detect a giant planet at 5 AU, implying that such planets are not rare. The properties of this Jupiter analog may be compared directly to those of the Jovian planets in our solar system. Its eccentricity is modest, e … 0:16, compared with e … 0:05 for both Jupiter and Saturn. Its mass is at least 4.0 MJUP (M sini). The two planets do not perturb each other signiicantly. Moreover, a third planet of sub-Jupiter mass could easily survive between these two known planets. Indeed, a third periodicity remains in the velocity measurements with P … 44:3 days and a semiamplitude of 13 m s 1 . This periodicity is caused either by a third planet at a … 0:24 AU or by inhomogeneities on the stellar surface that rotate with period 42 days. The planet interpretation is more likely, as the stellar surface is quiet both chromospherically [log﷿R 0 fi… 5 :0] and photospherically (brightness variations less than 1 mmag). Moreover, any hypothetical surface inhomogeneity would have to persist in longitude for 14 yr. Even with all three planets, an additional planet of terrestrial mass could orbit stably at 1 AU. The star 55 Cancri is apparently a normal, middle-aged main-sequence star with a mass of 0.95 M , rich in heavy elements (‰Fe=H …˛ 0 :27). This high metallicityraises the issue of the precise relationship between its age, rotation, and chromosphere. Subject headings: planetary systems N stars:individual (55 Cancri,HIP 43587,HD 75732, HR 3522, 1 Cancri)

Managing the paradox of inter‐firm learning: the role of governance mechanisms

Abstract: Organizational learning in inter‐firm exchange relationships poses a double‐edged sword. On one hand, inter‐firm learning is a desirable extension of organizational learning, developing a firm’s knowledge base, and providing fresh insights into strategies, markets, and relationships. On the other hand, inter‐firm learning can lead to unintended and undesirable skills transfer, resulting in the potential dilution of competitive advantage. This risk can be exacerbated by disparities in inter‐firm learning, resulting in uneven distribution of benefits and risks in the collaborative relationship. This paper articulates these two different views on inter‐firm learning, and second, develops a framework for the role of governance in regulating knowledge transfer. In particular, appropriate governance mechanisms must be crafted which match the learning intentions of the partners, the type of knowledge sought, and the designed duration for the collaboration, so as to maximize the benefits of learning while minimizing the risks. Implications for strategy and future research are offered.

The cell wall-associated kinase (WAK) and WAK-like kinase gene family.

Abstract: We have identified a large family of genes with sequence similarity to the cell wall-associated kinase ( WAK ) genes ([He et al., 1999][1]). Like the WAK s, these genes exist in multiple gene clusters, and our analyses suggest that they encode functional protein kinases that are associated with the

Timing of Entry in International Market: An Empirical Study of U.S. Fortune 500 Firms in China

Abstract: This study attempts to explain the timing of entry of firms in international markets. Based on the existing literature, we propose a framework that consists of firm-specific factors, industry/market factors, and host country factors. Empirical results, based on the entry information of U.S. Fortune 500 firms in China between 1979 and 1996, show that larger firms with greater level of internalization and scope economies are likely to enter this foreign market earlier. In addition, non-equity modes, competitors' behavior in the product market, and lower levels of country risk are significantly associated with early entry.

Differential resource use by primates and hornbills: implications for seed dispersal

Abstract: Arboreal frugivores, such as primates and hornbills, are important seed dis- persers for many tropical plant species, yet the degree to which they use the same resources is unknown. If primates and hornbills consume the same fruit species, they may be redundant in their roles as seed dispersers, and the loss of one of these taxa may be compensated for by the other. To examine resource use by tropical frugivores, we quantified the feeding habits of two hornbill species, Ceratogymna atrata and C. cylindricus, and five primate species, Colobus guereza, Lophocebus albigena, Cercopithecus pogonias, C. cephus, and C. nictitans, in the lowland rainforest of south-central Cameroon. Based on over 2200 feeding observations recorded between January and December 1998, we characterized the diets and estimated dietary overlap among frugivore species. Previous studies have cal- culated dietary overlap by counting the number of diet species that two animals share, often leading to inflated estimates of overlap. Our method incorporated the proportional use of diet species and fruit availability into randomization procedures, allowing a clearer as- sessment of the actual degree of overlap. This added complexity of analysis revealed that, although the diets of a hornbill and a primate species may have as many as 36 plant species in common, actual dietary overlap is low. These results suggested that there are distinct hornbill and primate feeding assemblages, with primates consuming a greater diversity of plant species and higher levels of nonfruit items like leaves and seeds. Using Correspon- dence Analysis, we also identified two primate assemblages, separated largely by degree of frugivory and folivory. In addition, we found that hornbills feed at significantly higher strata in the forest canopy and eat fruits of different colors than primates. Averaged across the year, overlap between groups (hornbill-primate) was significantly lower than combined within-group overlap (primate-primate and hornbill-hornbill), showing that primates and hornbills have dissimilar diets and are not redundant as seed dispersers. In equatorial Africa, primate populations face greater declines than hornbill populations because of hunting. It is unlikely that seed dispersal by hornbills will compensate for the loss of primates in maintaining forest structure.

Spatial and Temporal Association of As and Fe Species on Aquatic Plant Roots

Abstract: The formation of an Fe(III) precipitate (plaque) on the surface of aquatic plant roots may provide a means of attenuation and external exclusion of metals. Presently, the mechanisms of metal(loid) sequestration at the root surface are unresolved. Accordingly, we investigated the mechanisms of Fe and As attenuation and association on the roots of two common aquatic plant species, Phalaris arundinacea (reed canarygrass) and Typha latifolia (cattail) using X-ray absorption spectroscopy and X-ray fluorescence microtomography. Iron plaque of both P. arundinacea and T. latifolia consist predominantly of hydrated iron oxides (ferrihydrite) with lesser amounts of goethite and minor levels of siderite. Typha latifolia, however, differs from P. arundinacea by having a significant contribution from lepidocrocite as well as a greater proportion of crystalline minerals. Coexistence of goethite and lepidocrocite suggests the presence of chemically diverse microenvironments at the root surface. Arsenic exists as a combination of two sorbed As species, being comprised predominantly of arsenate- (approximately 82%) with lesser amounts (approximately 18%) of As(III)-iron (hydr)oxide complexes. Furthermore, both spatial and temporal correlations between As and Fe on the root surfaces were observed. While the iron (hydr)oxide deposits form a continuous surficial rind around the root, As exists in isolated regions on the exterior and interior of the root. Root surface-associated As generally corresponds to regions of enhanced Fe levels and may therefore occur as a direct consequence of Fe phase heterogeneity and preferential As sorption reactions.

Constraining the difference in convective blueshift between the components of alpha Centauri with precise radial velocities

Abstract: New radial velocities of Cen A & B obtained in the framework the Anglo-Australian Planet Search programme as well as in the CORALIE programme are added to those by Endl et al. (2001) to improve the pre- cision of the orbital parameters. The resulting masses are 1:105 0:0070 M and 0:934 0:0061 M for A and B respectively. The factors limiting how accurately these masses can be derived from a combined visual-spectroscopic solution are investigated. The total eect of the convective blueshift and the gravitational redshift is also inves- tigated and estimated to dier by 215 8m s 1 between the components. This suggests that the dierence in convective blueshift between the components is much smaller than predicted from current hydrodynamical model atmosphere calculations.

Bosnian Refugees and the Stressors of Exile: A Narrative Study

Abstract: The authors used semistructured interviews to examine exile-related stressors affecting a sample of 28 adult Bosnian refugees in Chicago. The interviews covered 3 areas: life in prewar Bosnia, the journey of exile, and, most centrally, life in Chicago. Primary sources of exile-related distress included social isolation and the loss of community, separation from family members, the loss of important life projects, a lack of environmental mastery, poverty and related stressors such as inadequate housing, and the loss of valued social roles. The implications of these findings for mental health interventions with refugees are considered, and the value of narrative methods in research with refugee communities is discussed.