Showing papers in "Zootaxa in 2020"

Catalog of the Biting Midges of the World (Diptera: Ceratopogonidae).

Abstract: A list of all valid 6,206 extant and 296 fossil species of Ceratopogonidae described worldwide is provided, along with all their synonyms. A full citation and the country of origin of the type is given, with some larger countries also providing a more specific state or province. For the first time, worldwide, nomina dubia are identified. Numbers of species of each genus and subgenus are listed. Within subfamilies and tribes, genera are listed alphabetically. Five species have newly recognized authors, four have new names and 28 new combinations are recognized, with these listed in a table. A commentary on the state of the systematics in the family and particularly of Culicoides Latreille is given. The museums of the world are listed with the types of various authors of Ceratopogonidae species indicated. Authors providing regional catalogs, as well as summation of various collections are tabulated. The rate of description since 1758 indicates a steady progression of description, with, for example, 1,231 valid species described since the compilation of the world species by Borkent Wirth (1997), till the end of 2018. The diversity in each Region is compared and the numbers of species shared between adjacent Regions presented.

Ixodidae (Acari: Ixodoidea): descriptions and redescriptions of all known species from 1758 to December 31, 2019

Abstract: References to the descriptions and redescriptions of the 742 species of Ixodidae published from 1758 to December 31, 2019 are compiled, with the goal of enabling tick taxonomists to readily access this diffuse and often confusing literature. Additionally, data resulting from this effort are critically analyzed to demonstrate the problems attending correct identification of several tick species that are of medical, veterinary and/or evolutionary importance, and to highlight the need for new or enhanced diagnostic techniques. Recent morphological and molecular studies indicate that some ixodid species names represent more than one taxon; therefore, it is expected that new species will be described in the near future, based partly on material already deposited in museums around the world.

Freshwater lampreys and fishes of Turkey; a revised and updated annotated checklist 2020

Abstract: The current status of the inland waters ichthyofauna of Turkey is revised, and an updated checklist of the freshwater fishes is presented. The latest checklist included all species listed in the available previous study that was published in 2015, which is now updated after a period of five years. We revised the validity of previously accepted species and added newly described and reported species in Turkey. Some previously erroneously reported species and not established alien fishes were excluded from this checklist. A total of 384 fish species belonging to 20 orders and 34 families have been reported in the inland waters of Turkey. Among these, 15 species (3.9%) are non-native and 208 species (54.2%) are considered as endemic to Turkey. A total of 119 species previously reported from Turkey have been excluded from Turkish ichthyofauna list, either in the present study or in previous studies.

Revisited morphology applied for Systematics of flat wasps (Hymenoptera, Bethylidae).

Abstract: The world fauna of the flat wasps (Bethylidae) is represented by about 3,000 valid species. The skeletal morphology of bethylids is still not adequately understood and the terminology is generally not standardized between its internal taxa and with other Hymenoptera families. The same scenario exists in most of the families in this order. To address this problem, we describe the external skeletal morphology of Bethylidae. We review the terms used to describe skeletal features in the Hymenoptera in general and a consensus terminology is proposed for Bethylidae, which is linked to the online Hymenoptera Anatomy Ontology. The morphology of the studied specimens is illustrated with photos and line drawings. We also discuss the morphological variation at both subfamilial and generic ranks. Our analyses challenge hundreds of inappropriate, confused or imprecise terms traditionally used for Hymenoptera morphology. As a result, we have applied hundreds of updates of the terminology available online at the Hymenoptera Anatomy Ontology.

Checklist of the dragonflies and damselflies (Insecta: Odonata) of Bangladesh, Bhutan, India, Nepal, Pakistan and Sri Lanka.

Abstract: A checklist of the dragonflies and damselflies occurring in Bangladesh, Bhutan, India (including Andaman and Nicobar Islands), Nepal, Pakistan and Sri Lanka is presented. In total 588 (including 559 full species) taxa are known to occur in the region of which 251 taxa (species & subspecies) are single country endemics. Recent taxonomic changes relevant to the area are summarized. Sixteen taxa are synonymized and a checklist of all synonyms established since 1950 is provided. Information is given on available larval descriptions including a list of genera present in the region for which no larvae have yet been described. Numerous species occurring in the area are still poorly known and a list of genera for which a revision is urgently needed is provided. The following new synonyms are established: Calicnemia sudhaae Mitra, 1994 = Calicnemia imitans Lieftinck, 1948 syn. nov.; Ceriagrion fallax cerinomelas Lieftinck, 1927 = Ceriagrion fallax Ris, 1914 syn. nov.; Ceriagrion fallax pendleburyi Laidlaw, 1931 = Ceriagrion fallax Ris, 1914 syn. nov.; Coenagrion kashmirus Chowdhary & Das, 1975 = Ischnura forcipata Morton, 1907 syn. nov.; Enallagma insula Fraser, 1920 = Aciagrion occidentale Laidlaw, 1919 syn. nov.; Himalagrion pithoragarhicum Sahni, 1964 = Ceriagrion fallax Ris, 1914 syn. nov.; Ischnura bhimtalensis Sahni, 1965 = Ischnura rubilio Selys, 1876 syn. nov.; Onychargia indica Sahni, 1964 = Paracercion calamorum (Ris, 1916) syn. nov.; Anaciaeschna kashmirense Singh & Baijal, 1954 = Anaciaeschna martini (Selys, 1897) syn. nov.; Cyclogomphus vesiculosus Selys, 1854 = Cyclogomphus ypsilon Selys, 1954 syn. nov.; Chlorogomphus brittoi Navas, 1934 = Chlorogomphus xanthoptera (Fraser, 1919) syn. nov.; Hylaeothemis indica Fraser, 1946 = Hylaeothemis apicalis Fraser, 1924 syn. nov.; Sympetrum durum Bartenef, 1916 = Sympetrum striolatum commixtum Selys, 1884 syn. nov.; Sympetrum himalayanum Navas, 1934 = Sympetrum hypomelas (Selys, 1884) syn. nov.; Sympetrum haematoneura Fraser, 1924 = Sympetrum speciosum Oguma, 1915 syn. nov.; Sympetrum speciosum taiwanum Asahina, 1951 = Sympetrum speciosum Oguma, 1915 syn. nov. In addition, Periaeschna lebasi Navas, 1930 is regarded a nomen nudum. The following new combinations are proposed: Onychogomphus acinaces Laidlaw, 1922 is transferred to the genus Melligomphus Chao, 1990 resulting in Melligomphus acinaces (Laidlaw, 1922) comb. nov. Onychogomphus saundersii Selys, 1854 is transferred to the genus Nychogomphus Carle, 1986 resulting in Nychogomphus saundersii (Selys, 1854) comb. nov..

Updated list of Anopheles species (Diptera: Culicidae) by country in the Afrotropical Region and associated islands.

Abstract: The distributions of the Afrotropical Anopheles mosquitoes were first summarized in 1938. In 2017, an extensive geo-coded inventory was published for 48 sub-Saharan African countries, including information such as sampling methods, collection dates, geographic co-ordinates and the literature consulted to produce the database. Using the information from the 2017 inventory, earlier distribution lists, museum collections and publications since 2016, this paper presents an updated, simplified list of Anopheles species by mainland countries and associated Afrotropical islands, with comments where applicable. It is intended as a supplement to the 2017 geo-coded inventory.

New species of Eurythenes from hadal depths of the Mariana Trench, Pacific Ocean (Crustacea: Amphipoda)

Abstract: Eurythenes S. I. Smith in Scudder, 1882 are one of the largest scavenging deep-sea amphipods (max. 154 mm) and are found in every ocean across an extensive bathymetric range from the shallow polar waters to hadal depths. Recent systematic studies of the genus have illuminated a cryptic species complex and highlighted the benefits of using a combination of morphological and molecular identification approaches. In this study, we present the ninth species, Eurythenes plasticus sp. nov., which was recovered using baited traps between the depths 6010 and 6949 m in the Mariana Trench (Northwest Pacific Ocean) in 2014. This new Eurythenes species was found to have distinct morphological characteristics and be a well-supported clade based on sequence variation at two mitochondrial regions (16S rDNA and COI). While this species is new to science and lives in the remote hadal zone, it is not exempt from the impacts of anthropogenic pollution. Indeed, one individual was found to have a microplastic fibre, 83.74% similar to polyethylene terephthalate (PET), in its hindgut. As this species has a bathymetric range spanning from abyssal to hadal depths in the Central Pacific Ocean basin, it offers further insights into the biogeography of Eurythenes.

The wild bees (Hymenoptera: Apoidea) of Morocco.

Abstract: Morocco is a well known hot-spot of biodiversity in the Mediterranean basin. While some taxa like vascular plants are relatively well recorded, important groups of pollinators like bees are still understudied. This article presents an updated checklist of the bee species of Morocco and includes a summary of global and regional distribution of each species. A total of 961 species belonging to six bee families and 68 genera are recorded: Andrenidae (8 genera, 217 species); Apidae (15 genera, 241 species); Colletidae (2 genera, 74 species), Halictidae (12 genera, 144 species), Megachilidae (28 genera, 271 species) and Melittidae (3 genera, 14 species). Among them, 67 species are recorded for the first time in Morocco. Around 70% of the bee fauna of Morocco consists of widespread Palaearctic species. Only 18% of Moroccan species recorded are restricted to North Africa and 8% are Moroccan single-country endemics (81 species). Afrotropical elements in the Moroccan fauna are few, with only 3% of Morocco species co-occuring in that region. This checklist is intended to stimulate new regional research on bees including their taxonomy and biogeography. As many groups of bees have been understudied, discovery of new species for science and new records for the country can be expected. Additional research including inventorying, monitoring, and integrative taxonomic studies are needed to develop a comprehensive strategy for bee conservation in Morocco.

A review of the genus Hemidactylus Goldfuss, 1820 (Squamata: Gekkonidae) from Angola, with the description of two new species.

Abstract: The genus Hemidactylus in Angola is represented by six species, all of them part of taxonomically and nomenclaturally challenging species complexes. We present a detailed taxonomic revision of the group in the region and describe two new species, Hemidactylus nzingae sp. nov. and Hemidactylus paivae sp. nov., both occuring in and potentially endemic to the highlands of Angola. Phylogenetic analysis using a combination of mitochondrial (ND2) and nuclear (MXRA5, PDC, RAG1) markers, as well as morphological and scalation data support the recognition of the new species. In addition, data support the revalidation of Hemidactylus bayonii Bocage, 1893, and Hemidactylus benguellensis Bocage, 1893. We also provide a redefinition of Hemidactylus longicephalus Bocage, 1873 with which we synonymize Hemidactylus mabouia molleri Bedriaga, 1892, from Sao Tome in the Gulf of Guinea. Given that the type material of H. bayonii, H. benguellensis, H. longicephalus and H. mabouia molleri have all been lost or destroyed, we designate neotypes for all of these nomina for purposes of nomenclatural stability. The description of the new species and the revision and revalidation of the Angolan species already described contributes to a better understanding of the taxonomy and biogeography of West and Central African Hemidactylus, as well as to the general biogeographic and evolutionary patterns of Angolan fauna. A key to the Angolan species is also presented.

A proposal for a new generic structure of the killifish family Aphaniidae, with the description of Aphaniops teimorii (Teleostei: Cyprinodontiformes).

Abstract: Following an exhaustive review of published molecular data and morphological characters, we propose a new generic classification for the Cyprinodontiform family Aphaniidae. The generic concept applied is based on monophyly, reasonable compactness, and morphological diagnoses. The proposed genera are monophyletic and compact groups that can be diagnosed by a combination of morphological characters. All species in Aphaniidae are morphologically homogeneous and have similar scale patterns, fin positions, and meristic characters. However, only a handful of morphological characters, including colour patterns, permit identification. We propose separation of Aphaniidae into eight monophyletic genera: Anatolichthys, Aphaniops, Aphanius, Kosswigichthys, Paraphanius, and Tellia, in addition to the new genera Esmaeilius and Apricaphanius, which are described for the species of the A. sophiae and A. iberus species groups, respectively. The original description of Aphanius hormuzensis does not fulfil the criteria of the International Commission on Zoological Nomenclature, therefore this taxon is re-described as A. teimorii. Esmaeilius arakensis, E. kavirensis, E. mesopotamicus, and E. pluristriatus are treated as synonyms of E. sophiae. Esmaeilius farsicus is a junior synonym of E. persicus (Jenkins, 1910). The fossil genus Brachylebias is considered as incertae sedis since it lacks diagnostic characters which would permit it to be assigned to any of the extant genera recognised in this study.

Accessing cryptic diversity in Neotropical rattlesnakes (Serpentes: Viperidae: Crotalus ) with the description of two new species

Abstract: Members of the Crotalus durissus species complex are widely distributed from Mexico to Argentina in areas with mainly seasonally dry tropical deciduous forest. Although four species (C. culminatus, C. durissus, C. simus and C. tzabcan) are currently recognized, species limits remain to be tested. Previous genetic studies suggest that C. durissus and C. simus may be paraphyletic and that at least one cryptic species may be present. We analyzed 2596 bp of DNA sequence data from three mitochondrial and one nuclear gene to infer phylogenetic relationships in the Neotropical rattlesnakes. We also examined museum and wild specimens as well as captive animals to analyze morphological characters. Our results suggest that the current taxonomy of the Crotalus durissus species complex does not reflect evolutionary history. We found strong support for five independent lineages within Crotalus simus (sensu lato), with genetic and morphological evidence for three previously recognized taxa and two new species, as well as three major lineages within C. durissus that each represent species hypothesis to be tested with additional evidence. We also found support to retain C. totonacus in the Crotalus molossus species complex. We suggest conservative taxonomic changes to the complex and related species, but more evidence is needed (e.g., morphology, ecology and venom composition) to clarify relationships among species.

Freshwater fishes from Paraná State, Brazil: an annotated list, with comments on biogeographic patterns, threats, and future perspectives.

Abstract: An annotated checklist for the freshwater fishes from Parana State, Brazil is provided. A total of 440 freshwater fish species are recorded for the state, distributed across five ecoregions: Upper Parana, Lower Parana, and Iguassu, all within the rio Parana basin, and corresponding to the state's Inland Slope, and Southeastern Mata Atlantica and Ribeira de Iguape, corresponding to the Atlantic Slope, encompassing minor coastal drainages emptying in the Baia de Paranagua or in the Baia de Guaratuba, and the rio Ribeira de Iguape basin, respectively. The Upper Parana ecoregion ocuppies the larger in area in the state, and is divided into the following sub-ecoregions: Floodplain, Paranapanema, Piquiri, and Ivai. Species richness for each ecoregion is as follows: 273 species for the Upper Parana (Paranapanema sub-ecoregion, 217 species; Floodplain sub-ecoregion, 193 species; Piquiri sub-ecoregion, 154 species; Ivai sub-ecoregion, 132 species), 154 species in the Lower Parana, 127 species in the Iguassu, 68 species in the Southeastern Mata Atlantica, and 50 species in the Ribeira de Iguape. We recorded 42 putatively undescribed species and 117 endemic species from specific ecoregions (except Upper Parana) or sub-ecoregions in the state. Ninety-eight species recorded are non-native from at least one of the state's ecoregions. Thirty-three species are considered threatened. The ecoregions in the Atlantic Slope share many more species with each other than with ecoregions in the Inland Slope. The Iguassu ecoregion is the only one located in Inland Slope that shares more species with the Atlantic Slope than with the remaining ecoregions from the Inland Slope. The Ivai sub-ecoregion lacks several species that are common to all other sub-ecoregions of the Upper Parana ecoregion. Comments on the historical development of taxonomic knowledge, biogeography, threats, and conservation strategies for the fish fauna from the Parana State are provided.

First Bocchus Ashmead (Hymenoptera: Dryinidae) from Upper Eocene Rovno amber: B. schmalhauseni sp. nov.

Abstract: A new species of pincer wasps, Bocchus schmalhauseni sp. nov. (Dryinidae) is described from Rovno amber (Ukraine). The amber piece was collected in the Vladimirets District in the north-west of the Rovno region. Taking into account this new record, a total of five species of Dryinidae are registered from Rovno amber. A key to the fossil species of Bocchus Ashmead is presented. The possible reasons for the different distribution of Dryinus species on both coasts of Subparathetys are discussed.

Checklist, Bibliography and Quantitative Data of the Arthropods of Hispaniola

Abstract: An updated and extensively revised checklist of the arthropods of Hispaniola (Dominican Republic and Haiti) is presented 11 years after the publication of the original in 2008. It integrates and quantifies all the terrestrial and surrounding marine arthropod species (plus those of Tardigrada and Onychophora), reported in the zoological literature for Hispaniola through the middle of 2019. A total of 9,920 valid species (8,202 extant and 1,718 fossil) are listed, which represents an increase of 1,683 species (1,369 extant and 314 fossil) from the original list. The largest component is Insecta (6,784 extant and 1,136 fossil), including 2,206 extant species of Coleoptera, 1,042 species of Hemiptera, 929 species of Diptera, 913 species of Lepidoptera and 774 species of Hymenoptera. Emphasis is on reviewing and updating the original list, including all newly recorded taxa and all pertinent taxonomic changes proposed since then. Important corrections have been made, and explanatory notes have been added. For example, multiple authors have confused the Lesser Antillean island of Dominica with the Dominican Republic. This error is much more common in the literature than was initially recognized. Erroneous records attributing species from one island to the other have been identified and corrected. The original spelling of the cricket species Scapsipedus bastardoi Otte Perez-Gelabert, 2009, dedicated to Dominican biologist Ruth H. Bastardo, is corrected to Scapsipedus bastardoae nom. emend. High species endemism is typical of the biota of Caribbean islands. In this checklist, a total of 3,161 arthropod species (38.6%) are considered endemic or unique to Hispaniola. Among the speciose groups with higher levels of endemism are the Diplopoda (91.6%), Orthoptera (90.1%), Trichoptera (82.6%), Coleoptera (49.3%) and Araneae (47.5%). Also, a total of 201 arthropod species (174 insects + 27 non-insects) are identified as introduced to the island. The accompanying bibliography complements the taxonomic information and includes over 5,000 titles.

Checklist of gobies (Teleostei: Gobiidae) of the Mediterranean Sea and a key for species identification.

Abstract: A checklist of 73 gobiid species (Teleostei: Gobiidae) recorded to date from the Mediterranean Sea is established following the evidence approach for checklists. The Mediterranean gobiofauna currently has 62 known native species and 11 alien species. An identification key to gobiid species known from the area is provided. The principles of character selection for the key are discussed.

Disentangling vines: a study of morphological crypsis and genetic divergence in vine snakes (Squamata: Colubridae: Ahaetulla) with the description of five new species from Peninsular India.

Abstract: We carried out a taxonomic revision of Ahaetulla species inhabiting Peninsular India, using a multiple criteria approach (including genetics, morphology, and geography). Our work included populations of the A. nasuta complex (widespread across the entire region, including the Western Ghats), the A. pulverulenta complex (in the Western Ghats, within Peninsular India) and the A. dispar complex (endemic to the Southern Western Ghats) which all revealed undocumented cryptic diversity. Here, we describe five new species and effect nomenclatural changes to some recognised taxa. In the A. nasuta complex, we describe four species from several latitudinal blocks of the Western Ghats and make nomenclatural emendations to the plains populations in the Indian peninsula. We effect nomenclatural change in the A. pulverulenta population of the Western Ghats and describe a new species from the A. dispar group. Our study highlights the use of a multi-criteria approach in unraveling cryptic diversity. This study also reveals a striking case of discordance between morphological and genetic divergence, and the way this is reflected in previous taxonomic and nomenclatural treatments of these populations.

A review of the African snake-eyed skinks (Scincidae: Panaspis) from Angola, with the description of a new species.

Abstract: The genus Panaspis in Angola is represented by four species, most of them part of taxonomically and nomenclaturally challenging species-complexes. We present a taxonomic revision of the group in the region and describe one new species, Panaspis mocamedensis sp. nov., endemic to the lowland areas of the Namibe province, southwestern Angola. Phylogenetic analysis using a combination of mitochondrial (16S, cytb) and nucleares (RAG1, PDC) markers, as well as morphological and meristic data support the recognition of the new species. In addition, these data support the presence of nominotypical Panaspis cabindae, P. wahlbergi and P. maculicollis in Angola. Reexamination of the Angolan population of P. breviceps was based on morphological analysis, as no molecular data from Angola is available for this species. According to our results, this population likely represents the nominotypical form, but due to its complex taxonomic and nomenclatural history and the lack of molecular data, this population needs to be reconsidered when molecular data become available. The description of a new species and revision of the Angolan Panaspis contributes to a better understanding of the true species richness of the Angolan herpetofauna, as well as to understanding the major biogeographic patterns of the region. A key to Angolan Panaspis species is also presented.

Checklist of decapod crustaceans from the coast of the São Paulo state (Brazil) supported by integrative molecular and morphological data: III. Infraorder Brachyura Latreille, 1802

Abstract: This checklist is the third part of a series derived from a long-term multidisciplinary project on the biodiversity of decapod crustaceans from marine and coastal environments of Sao Paulo state (Brazil). We integrated molecular techniques (DNA markers) and morphological analyses of adult specimens for accurate identifications. We compilated 185 species from the literature, but we confirmed the presence of 168 species: 130 of which we sampled, analyzed and obtained sequences (COI and/or 16S totalizing 113 sequences) and 38 that were not directly collected but were confirmed by analyses. In addition, 27 had their distribution removed from Sao Paulo due to uncertainties, and absence of material as voucher. Five species were reported for the first time on the coast of Sao Paulo (Calappa ocellata, Neohelice granulata, Teleophrys pococki, Teramnonotus monodi, Tetraxanthus rathbunae) and one on the Brazilian coast (Pseudomedaeus agassizi). Most of the non-sampled species previously reported on the coast of Sao Paulo might be considered doubtful records stablished in the past by inaccurate analyses, which assumed extended distribution to the area and/or misidentifications. At this time and based on our refined compilation, we can estimate the brachyuran diversity on the coast of Sao Paulo in 168 species. This detailed inventory contributes to the knowledge on the local decapod fauna by checking available dataset, adding new species records in Sao Paulo and new sequences to GenBank database. These data may serve as baseline for future identifications and studies on conservation, population genetics, biogeography and phylogenetics, which might flag species that deserve further investigations and concerns.

Bioregionalisation of the freshwater zoogeographical areas of mainland China

Abstract: Biogeographic regionalisations extract patterns of co-occurrence from different taxa to form a hierarchical system of geographical units of different scales. This system is useful for revealing biogeographic patterns and can be used as the basis for scientific communication between different fields. The history of Chinese freshwater biogeography is not well known to most modern biogeographers and is reviewed herein. We produce the first quantitative bioregionalisation of the freshwater zoogeographic areas of mainland China based on multiple animal groups. The combined occurrence data of amphibians, freshwater fish and freshwater crabs were subjected to cluster and network analyses. The two different methods yielded largely similar results. We propose four freshwater zoogeographical subregions (Beifang, Tarim, China, and the Tibetan subregion), three dominions for the China subregion (Jianghuai, Dongyang, and the new Dian dominion), three provinces for the Dian dominion (West Hengduan, Diannan Highlands and the new Yungui Plateau province) and two provinces for the Dongyang dominion (Zhemin and the new Huanan province) according to the naming rules of ICAN. The endemic areas of each animal group were then individually studied and were found to reflect the bioregionalisation at the subregion level, but differed from each other at the dominion and province level. Our analyses show that: (1) previous intuitive biogeographical studies have found similar areas; (2) there are recurring large scale biogeographic patterns in Chinese freshwater fishes, amphibians and freshwater crabs; and (3) bioregionalisations derived from quantitative methods can be effective for partitioning areas into biogeographically meaningful units.

A new bee-mimicking stiletto fly (Therevidae) from China discovered on iNaturalist.

Abstract: The Chinese stiletto fly fauna is poorly known, with few species previously described and no endemic genera. A new genus and species of charismatic stiletto fly is described from northern China. Although the species was previously known from a poorly preserved specimen, the true form of this apparent bumble bee-mimic was not known until images of a live individual were discovered amongst photographs posted on iNaturalist, an online citizen scientist and biodiversity social network.

Bark beetles and pinhole borers recently or newly introduced to France (Coleoptera: Curculionidae, Scolytinae and Platypodinae).

Abstract: We present an annotated list of 11 Scolytinae and Platypodinae species newly or recently introduced to France. Four species are recorded for the first time as interceptions: Euplatypus hintzi (Schaufuss), Euplatypus parallelus (Fabricius), Xyleborus affinis Eichhoff and Xyleborus ferrugineus (Fabricius). Two are possibly naturalised: Xyleborus bispinatus Eichhoff and Cryphalus dilutus Eichhoff, while Cyclorhipidion distinguendum (Eggers) and Xyloterinus politus (Say) are confirmed as species newly established in Europe. Moreover, an unidentified species of Amasa Lea, collected previously in Spain, is recorded for the first time in France: Amasa sp. near truncata (Erichson). We point out that literature references to Amasa truncata as an invasive species in New Zealand and South America are incorrect, as the photographs of these non-native populations do not match the holotype of A. truncata. For each species we have updated its global distribution, detailed all French records, and summarized biology, ecology, host trees and potential risks as pests.

The invertebrate host of salmonid fish parasites Ceratonova shasta and Parvicapsula minibicornis (Cnidaria: Myxozoa), is a novel fabriciid annelid, Manayunkia occidentalis sp. nov. (Sabellida: Fabriciidae).

Abstract: Myxosporea (Cnidaria: Myxozoa) are common fish parasites with complex life cycles that involve annelid hosts. Two economically important salmonid-infecting myxosporeans from rivers of the northwestern United States, Ceratonova shasta (Noble, 1950) and Parvicapsula minibicornis Kent et al., 1997, have life cycles that require a freshwater annelid host, identified previously as Manayunkia speciosa Leidy, 1859. This species was described originally from Pennsylvania, with subsequent records from New Jersey, the Great Lakes and west coast river basins. Despite apparent widespread distributions of both suitable fish hosts and the nominal annelid host, both parasites are restricted to river basins in the northwestern US and have never been recorded from the Great Lakes or the eastern US. In this study, we sampled 94 infected and uninfected annelids from two northwestern US rivers to confirm the identity of the host. We found these new specimens had mitochondrial COI sequences with no more than 4.5% distance from each other, but with at least 11% divergence from M. speciosa sampled from near the type locality (New Jersey) and Lake Superior. We did not recover any M. speciosa from either west coast river. The annelid from the Klamath and Willamette rivers showed marked sexual dimorphism that has not been reported in any Manayunkia described to date, though it is apparent that this had been missed in M. speciosa. Accordingly, we describe a new taxon, Manayunkia occidentalis sp. nov., and show that it can host both C. shasta and P. minibicornis. We suspect that previous records of Manayunkia from Pacific Northwest watersheds are likely to be M. occidentalis sp. nov. and not M. speciosa. Sampling of Manayunkia from additional localities is underway to test if the novel Manayunkia species is the only freshwater fabriciid annelid present across the Pacific Northwest.

Hydroids (Cnidaria, Hydrozoa) from Mauritanian Coral Mounds.

Abstract: Agassiz, L. (1862) Contributions to the natural history of the United States of America. Vol. 4. Little Brown, Boston, 380 pp.Aguirrezabalaga, F., Altuna, A., Borja, A., Feliu, J., GarcíaCarrascosa, A.M., Romero, A., San Vicente, C., Torres-Gómez-de-Cádiz, J.A., Uriz, M.J. Ibánez, M. (1984) Contribución al conocimiento de la fauna marina de la costa Vasca. Π. Lurralde, Investigación y espacio, 1984, pp. 83-133.Alder, J. (1856) A notice of some new genera and species of British hydroid zoophytes. Annals and Magazine of Natural History, Series 2, 18, 353-362. https://doi.org/10.1080/00222935608697652Allman, G.J. (1873) Interim report on the hydroids collected by L.F. de Pourtalès during the Gulf Stream exploration of United States coast survey. Bulletin of the Museum of Comparative Zoology at Harvard College, 3 (7), 185-186.Allman, G.J. (1874) Report on the Hydroida collected during the expeditions of H.M.S. 'Porcupine.' Transactions of the Zoological Society of London, 8, 469-481. https://doi.org/10.1111/j.1096-3642.1874.tb00566.xAllman, G.J. (1877) Report on the Hydroida collected during the exploration of the Gulf Stream by L.F. de Pourtalès, assistant United States Coast Survey. Memoirs of the Museum of Comparative Zoology at Harvard College, 5 (2), 1-66. https://doi.org/10.5962/bhl. TITLE 15852Allman, G.J. (1883) Report on the Hydroida dredged by H.M.S. Challenger during the years 1873-76. I. Plumularidae. Report of the Scientific Results of the Voyage of H.M.S. Challenger 1873-1876, 1-55, pls. 1-20. https://doi.org/10.5962/bhl. TITLE 11299Altuna, A. (2012) New records of bathyal Leptolida (Cnidaria: Hydrozoa: Leptothecata) from the Bay of Biscay and the northwestern Iberian Peninsula (northeastern Atlantic). Zootaxa, 3565 (1), 1-17. https://doi.org/10.11646/zootaxa.3565.1.1Altuna Prados, A. Álvarez Claudio, C. (1994) El género Zygophylax Quelch, 1885 (Cnidaria, Hydrozoa) en el Golfo de Vizcaya. Miscel.lània Zoològica, 17, 1-16.Alvarez Claudio, M.C. (1993) Hidrozoos bentonicos y Catálogo de Antozoos de la Plataforma y Talud Continentales de la costa central de Asturias. Tesis de Licenciatura, Universidad de Oviedo, Oviedo, Asturias, 458 pp.Alvarez Claudio, M.C. (1995) Some records of the superfamily Plumularioidea L. Agassiz, 1862 (Cnidaria, Hydrozoa) from the Bay of Biscay. Miscel·lània Zoològica, 18, 9-20.Ansín Agís, J. (1992) Hidrozoos de la Ría de Vigo. Tesis de Licenciatura, Universidad de Vigo, Ourense, Galicia, 282 pp.Ansín Agís, J. (1998) Plumularioidea (Cnidaria, Hydrozoa) recolectados por las expediciones holandesas "CANCAP" en el Atlántico nor-oriental. Tesis Doctoral, Universidade de Vigo, Vigo, Galicia, 675 pp. Ansín Agís, J., Ramil, F. Vervoort, W. (2001) Atlantic Leptolida (Hydrozoa, Cnidaria) of the families Aglaopheniidae, Halopterididae, Kirchenpaueriidae and Plumulariidae collected during the CANCAP and Mauritania-II expeditions of the National Museum of Natural History, Leiden, The Netherlands. Zoologische verhandelingen, Leiden, 333, 1-268.Ansín Agís, J., Vervoort, W. Ramil, F. (2014) Hydroids of the families Kirchenpaueriidae Stechow, 1921 and Plumulariidae McCrady, 1859 (Cnidaria, Hydrozoa) collected in the Western Pacific Ocean by various French Expeditions. Zoosystema, 36, 789-840. https://doi.org/10.5252/z2014n4a6Arévalo y Carretero, C. (1906) Contribución al estudio de los Hidrozoarios españoles existentes en la Estación de Biología marítima de Santander. Memorias de la Real Sociedad Española de Historia Natural, 4 (3), 79-109.Avila, S.P. Malaquias, M.A.E. (2003) Biogeographical relationships of the molluscan fauna of the Ormonde Seamount (Gorringe Bank, Northeast Atlantic Ocean). Journal of Molluscan Studies, 69, 145-150. https://doi.org/10.1093/mollus/69.2.145Bale, W.M. (1915) Report on the Hydroida collected in the Great Australian Bight and other localities. Part III. Fish. Zool. (biol) Biological Results of the Fishing Experiments Carried on by the F.I.S. "Endeavour", 1909-14, 3 (5), 304-307.Bedot, M. (1916) Sur le genre Kirchenpaueria. Revue Suisse de Zoologie, 24 (11), 637-648.Bedot, M. (1917) Le genre Nemertesia. Mémoire de la Société de Physique et d'Histoire Naturelle de Genève, 39 (1), 15-52.Bedot, M. (1921a) Notes systématiques sur les plumularides. 2me partie. Revue Suisse de Zoologie, 29 (1), 1-40. https://doi.org/10.5962/bhl.part.84689Bedot, M. (1921b) Hydroïdes provenant des campagnes des yachts Hirondelle et Princesse-Alice (1887-1912). I. Plumulariidae. Résultats des Campagnes scientifiques accomplies sur son yacht par le Prince Albert Ier de Monaco, 60, 1-73.Bedot, M. (1923) Notes systématiques sur les plumularides. 3me partie. Revue Suisse de Zoologie, 30 (7), 213-243.Billard, A. (1901) Note sur l'Antennularia antennina Lin. et sur l'A. perrieri n. sp. Bulletin du Museìum national d'histoire naturelle, Paris, 7, 68-75.Billard, A. (1904) Contribution à l'étude des Hydroïdes (multiplication, regeneration, greffes, variations). Annales des sciences naturelles, Zoologie, Series 8, 20, 1-251.Billard, A (1906a) Hydroïdes. In: Mission des pêcheries de la côte occidental d'Afrique, III. Actes de la Société Linnéenne de Bordeaux, 61, 69-76.Billard, A. (1906b) Hydroïdes. In: Expéditions scientifiques du "Travailleur" et du "Talisman" pendant les années 1880, 1881, 1882, 1883, etc. Masson Cie., Paris, pp. 153-243.Billard, A. (1906c) Note sur les Hydroïdes du Travailleur et du Talisman. Bulletin du Museìum national d'histoire naturelle, 12 (5), 329-334.Billard, A. (1931) Hydroïdes de Mauritanie. Bulletin du Muséum national d'histoire naturelle, Series 2, 3 (7), 673-678.Billard, A. (1934) Note sur quelques hydroïdes du Maroc. Bulletin de la Société Zoologique de France, 59, 227-231. Boero, F. Bouillon, J. (1993) Zoogeography and life cycle patterns of Mediterranean hydromedusae (Cnidaria). Biological Journal of the Linnean Society, 48, 239-266. https://doi.org/10.1111/j.1095-8312.1993.tb00890.xBoero, F. Fresi, E. (1986) Zonation and evolution of a rocky bottom hydroid community. Marine Ecology, 7 (2), 123-150. https://doi.org/10.1111/j.1439-0485.1986.tb00152.xBouillon, J. (1984) Révision de la famille des Phialuciidae (Kramp, 1955) (Leptomedusae, Hydrozoa, Cnidaria), avec un essai de classification des Thecatae-Leptomedusae. Indo-Malayan Zoology, 1, 1-24.Bouillon, J., Massin, C. Kresevic, R. (1995) Hydroidomedusae de l'Institut royal des Sciences naturelles de Belgique. Documents de travail de l'Institut royal des Sciences naturelles de Belgique, 78, 3-106.Bouillon, J., Medel, M.D., Pagès, F., Gili, J.M., Boero, F. Gravili, C. (2004) Fauna of the Mediterranean Hydrozoa. Scientia Marina, 68 (2), 5-438. https://doi.org/10.3989/scimar.2004.68s25Bouillon, J., Gravili, C., Pagès, F., Gili, J.M. Boero, F. (2006) An introduction to Hydrozoa. Editions du Muséum, Paris, 591 pp.Broch, H. (1913) Hydroida from the "Michael Sars" North Atlantic Deep-Sea Expedition 1910. In: Report on the scientific results of the "Michael Sars" north Atlantic deep-sea expedition 1910 3(1) Zoology, 1-18.Broch, H. (1914) Hydrozoa benthonica. In: Michaelsen, W. (Ed.), Beiträge zur Kenntnis der Meeresfauna Westafrikas. Vol. 1. Friederichsen, Hamburg, pp. 19-50.Browne, E.T. (1907) The Hydroida collected by the "Huxley" from the North Side of the Bay of Biscay in August, 1906. Journal of the Marine Biological Association of the United Kingdom, 8, 15-37. https://doi.org/10.1017/S002531540004371XBuchanan, J.B. (1957) The hydroid fauna of the Gold Coast. Revue de Zoologie et de Botanique Africaines, 56 (3-4), 349-372.Burdon-Jones, C. Tambs-Lyche, H. (1960) Observations on the fauna of the North Brattholmen stone-coral reef near Bergen. Årbok for Universitetet i Bergen, Matematisk-naturvitenskapling, 1960 (4), 1-24.Cairns, S.D. Chapman, R.E. (2001) Biogeographic affinities of the North Atlantic deep-water Scleractinia. In: Willison, J.H.M., Hall, J., Gass, S.E., Kenchington, E.L.R., Butler, M. Doherty, P. (Eds.), Proceedings of the First International Sympo-sium on Deep-Sea Corals. Ecology Action Centre, Halifax, pp. 30-57Calder, D.R. (1988) Shallow-water hydroids of Bermuda: the Athecatae. Royal Ontario Museum, Life Sciences Contributions, 148, 1-107. https://doi.org/10.5962/bhl. TITLE 52225Calder, D.R. (1991) Shallow-water hydroids of Bermuda. The Thecatae, exclusive of Plumularioidea. Royal Ontario Museum, Life Sciences Contributions, 154, 1-140.Calder, D.R. (1997) Shallow-water hydroids of Bermuda: superfamily Plumularioidea. Royal Ontario Museum, Life Sciences Contributions, 161, 1-85.Calder, D.R. (1998) Hydroid diversity and species composition along a gradient from shallow waters to deep sea around Bermuda. Deep-Sea Research I, 45, 1843-1860. https://doi.org/10.1016/S0967-0637(98)00044-2Calder, D.R. (2012) On a collection of hydroids (Cnidaria, Hydrozoa, Hydroidolina) from the west coast of Sweden, with a checklist of species from the region. Zootaxa, 3171 (1), 1-77. https://doi.org/10.11646/zootaxa.3171.1.1Calder, D.R. Vervoort, W. (1998) Some hydroids (Cnidaria: Hydrozoa) from the Mid-Atlantic Ridge, in the North Atlantic Ocean. Zoologische verhandelingen, Leiden, 319, 1-65.Calero, B., Ramil, F. Ramos, A. (2017) Echinoderms of the Mauritanian Deep-Sea Waters. In: Ramos, A., Ramil, F. Sanz, J.L. (Eds.), Deep sea ecosystems off Mauritania: Researching marine biodiversity and habitats in West African Deep-waters. Springer, Dordrecht, pp. 445-480. https://doi.org/10.1007/978-94-024-1023-5_12Castillo, S. (2017) Marine Mollusc (Gastropoda and Bivalvia) from Northwest Africa. Undergraduate dissertation, Universidad de Vigo, Pontevedra, Ourense, Vigo, Galicia, 422 pp.Christiansen, B.O. (1972). The hydroid fauna of the Oslo Fjord in Norway. Norwegian Journal of Zoology, 20, 279-310.Clarke, S.F. (1879) Report on the Hydroida collected during the Exploration of the Gulf Stream of Mexico by A

The marine ichthyofauna of Lebanon: an annotated checklist, history, biogeography, and conservation status.

Abstract: This is an annotated checklist of the marine fish species of Lebanon (Levant coast, eastern Mediterranean). It comprises a total of 367 fish species distributed in 159 families and 27 orders, out of which 70 species are non-indigenous and 28 are recorded for the first time from Lebanon. The checklist includes all names and citations of species found in the literature, as well as references to specimens available in museum collections. It also includes an updated zoogeographic distribution and conservation status for each species. Threats were estimated based on surveys and observations made over the last two decades; broadly categorized conservation actions were also suggested.

A revision of the Dwarf Geckos, genus Lygodactylus (Squamata: Gekkonidae), from Angola, with the description of three new species.

Abstract: At present the genus Lygodactylus is represented by three species in Angola confirmed by voucher specimens—L. angolensis, L. bradfieldi, and L. capensis—and two others believed to be present, but without specimens with precise localities, L. chobiensis and L. lawrencei. We present a detailed taxonomic revision of the group in Angola and describe three new species, Lygodactylus baptistai sp. nov. L. nyaneka sp. nov. and Lygodactylus tchokwe sp. nov. Phylogenetic analysis using the mitochondrial marker ND2, as well as morphological data support the recognition of the new species. In addition, data suggest that specimens historically assigned to L. capensis in Angola represent misidentifications of L. nyaneka sp. nov. and L. tchokwe sp. nov. We revisit the identity of Lygodactylus laurae, a junior synonym of L. angolensis. We also present the first confirmed record of L. lawrencei in the country, using both morphological and molecular data. The description of the new species and the revision of the taxonomic identity of the Angolan populations of the genus, raises the number of species occurring in the country to five. A key to the Angolan species is presented.

Unraveling the species diversity and relationships in the Leptodactylus mystaceus complex (Anura: Leptodactylidae), with the description of three new Brazilian species.

Abstract: Members of the Leptodactylus mystaceus species complex are widely distributed in forests and open formations of South America east of the Andes. Species of the complex are morphologically similar or indistinguishable among each other, but acoustic data have been the cornerstone for species discrimination across their geographic ranges. In this paper, we re-examine the monophyly, species diversity, and relationships in the L. mystaceus complex on the basis of morphology, coloration, acoustics, and DNA sequences. Morphological and color patterns originally used to the allocation of species to the L. mystaceus complex are also reassessed. Our results revealed three new species from the Brazilian Atlantic Forest and Cerrado, which are named and described herein, based mainly on acoustic and molecular data. Populations assigned to the lineage widely distributed across the South American Dry Diagonal (DD), reported in this study as L. cf. mystaceus, is likely paraphyletic with respect to the nominal species (Amazonian lineage), but additional data are still needed to address the taxonomic status of the DD lineage.

Additions to the taxonomy of Gnamptogenys Roger (Hymenoptera: Formicidae: Ectatomminae) with an updated key to the New World species

Abstract: This study aimed to raise and address nomenclatural issues surrounding Neotropical species in the genus Gnamptogenys, in addition to describing new species accumulated in myrmecological collections. New and reinstated names recognized here include G. pernambucana (Santschi) sp. rev., stat.n., G. lenis sp. n., G. latistriata sp. n., and G. avus sp. n. The queen and intercaste of G. lavra Lattke and the queen of G. pernambucana are described for the first time. For these five species we provide complete descriptions and diagnoses, comments on taxonomy and natural history, distribution data, and high-resolution images, including the first images of G. lavra. An updated identification key for all the Neotropical species of Gnamptogenys is also provided, including the new and revived species as well as G. aspera Morgan and G. pilosa Lattke, not present in previous keys for the genus.

Two new species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from a karstic archipelago in the Salween Basin of southern Myanmar (Burma).

Abstract: An integrative taxonomic analysis of the 10 species of the Cyrtodactylus sinyineensis group based on squamation, color pattern, and the mitochondrial gene NADH dehydrogenase subunit 2 (ND2) and its flanking tRNA regions, recovered the newly discovered populations from Datt Kyaik and Taung Wine Hills in Kayin State as the new species Cyrtodactylus dattkyaikensis sp. nov. and C. taungwineensis sp. nov. The Maximum Likelihood and Bayesian phylogenetic estimates supported C. dattkyaikensis sp. nov. as the sister species of C. bayinnyiensis and C. taungwineensis sp. nov. as the sister species of C. sinyineensis. Each new species is differentially diagnosable from all other C. sinyineensis group species based on their morphological placement in multivariate space and several statistically significant mean differences is meristic squamation and color pattern data. The C. sinyineensis group ranges across an archipelago of karstic habitat-islands in the Salween Basin of southern Myanmar. The discovery of these new species continues to underscore the unprecedented high degree of diversity and site-specific endemism in this relatively small region and the urgent need for the conservation of its karstic terranes.

Morphological and molecular description of a new genus and species of black coral (Cnidaria: Anthozoa: Hexacorallia: Antipatharia: Antipathidae: Blastopathes) from Papua New Guinea

Abstract: Blastopathes medusa gen. nov., sp. nov., is described from Kimbe Bay, Papua New Guinea, based on morphological and molecular data. Blastopathes, assigned to the Antipathidae, is a large, mythology-inspiring black coral characterized by clusters of elongate stem-like branches that extend out at their base and then curve upward. Colonies are not pinnulate and contain single branches, which could represent new branch cluster formations. Morphological and molecular (mitochondrial DNA and targeted capture of nuclear loci) evidence supporting the establishment of a new genus is discussed. This is the first study to utilize the target capture of ultraconserved elements (UCEs) and exonic loci to elucidate phylogenetic relationships among black corals and to identify and place a new genus and species.

Systematics of big-eyed bats, genus Chiroderma Peters, 1860 (Chiroptera: Phyllostomidae)

Abstract: We present a revision of the Neotropical bat genus Chiroderma, commonly known as big-eyed bats. Although species of Chiroderma have a wide distribution from western Mexico to southern Brazil, species limits within Chiroderma are not clearly defined, as attested by identification errors in the literature, and there is no comprehensive revision of the genus that includes morphological and molecular data. Our review is based on phylogenetic analyses of two mitochondrial (COI and CYTB) and two nuclear (RAG2 and DBY) genes, coalescence analyses of mitochondrial genes, and morphological analyses including type specimens of all named taxa. We recognize seven species in three clades: the first clade includes (1) C. scopaeum Handley, 1966, endemic to western Mexico and previously considered a subspecies of C. salvini; and (2) C. salvini Dobson, 1878, a taxon associated with montane forests, distributed from Mexico to Bolivia; the second clade includes (3) C. improvisum Baker and Genoways, 1976, endemic to the Lesser Antilles, and (4) C. villosum Peters, 1860, widely distributed on the continental mainland and polytypic, with subspecies C. v. villosum and C. v. jesupi; and the third clade includes (5) the polytypic C. doriae Thomas, 1891, with C. d. doriae distributed in eastern Brazil and Paraguay, and C. d. vizottoi, occurring in northeastern Brazil; (6) C. trinitatum Goodwin, 1958, distributed from Trinidad to Amazonia; and (7) C. gorgasi Handley, 1960, distributed from Honduras to trans-Andean South America, previously considered a subspecies of C. trinitatum.