We synthesize data on all known extant and fossil Coleoptera family-group names for the first time. A catalogue of 4887 family-group names (124 fossil, 4763 extant) based on 4707 distinct genera in Coleoptera is given. A total of 4492 names are available, 183 of which are permanently invalid because they are based on a preoccupied or a suppressed type genus. Names are listed in a classification framework. We recognize as valid 24 superfamilies, 211 families, 541 subfamilies, 1663 tribes and 740 subtribes. For each name, the original spelling, author, year of publication, page number, correct stem and type genus are included. The original spelling and availability of each name were checked from primary literature. A list of necessary changes due to Priority and Homonymy problems, and actions taken, is given. Current usage of names was conserved, whenever possible, to promote stability of the classification.

New synonymies (family-group names followed by genus-group names): Agronomina Gistel, 1848 syn. nov. of Amarina Zimmermann, 1832 (Carabidae), Hylepnigalioini Gistel, 1856 syn. nov. of Melandryini Leach, 1815 (Melandryidae), Polycystophoridae Gistel, 1856 syn. nov. of Malachiinae Fleming, 1821 (Melyridae), Sclerasteinae Gistel, 1856 syn. nov. of Ptilininae Shuckard, 1839 (Ptinidae), Phloeonomini Adam, 2001 syn. nov. of Omaliini MacLeay, 1825 (Staphylinidae), Sepedophilini Adam, 2001 syn. nov. of Tachyporini MacLeay, 1825 (Staphylinidae), Phibalini Gistel, 1856 syn. nov. of Cteniopodini Solier, 1835 (Tenebrionidae); Agronoma Gistel 1848 (type species Carabus familiaris Duftschmid, 1812, designated herein) syn. nov. of Amara Bonelli, 1810 (Carabidae), Hylepnigalio Gistel, 1856 (type species Chrysomela caraboides Linnaeus, 1760, by monotypy) syn. nov. of Melandrya Fabricius, 1801 (Melandryidae), Polycystophorus Gistel, 1856 (type species Cantharis aeneus Linnaeus, 1758, designated herein) syn. nov. of Malachius Fabricius, 1775 (Melyridae), Sclerastes Gistel, 1856 (type species Ptilinus costatus Gyllenhal, 1827, designated herein) syn. nov. of Ptilinus Geoffroy, 1762 (Ptinidae), Paniscus Gistel, 1848 (type species Scarabaeus fasciatus Linnaeus, 1758, designated herein) syn. nov. of Trichius Fabricius, 1775 (Scarabaeidae), Phibalus Gistel, 1856 (type species Chrysomela pubescens Linnaeus, 1758, by monotypy) syn. nov. of Omophlus Dejean, 1834 (Tenebrionidae). The following new replacement name is proposed: Gompeliina Bouchard, 2011 nom. nov. for Olotelina Baguena Corella, 1948 (Aderidae).

Reversal of Precedence (Article 23.9) is used to conserve usage of the following names (family-group names followed by genus-group names): Perigonini Horn, 1881 nom. protectum over Trechicini Bates, 1873 nom. oblitum (Carabidae), Anisodactylina Lacordaire, 1854 nom. protectum over Eurytrichina LeConte, 1848 nom. oblitum (Carabidae), Smicronychini Seidlitz, 1891 nom. protectum over Desmorini LeConte, 1876 nom. oblitum (Curculionidae), Bagoinae Thomson, 1859 nom. protectum over Lyprinae Gistel 1848 nom. oblitum (Curculionidae), Aterpina Lacordaire, 1863 nom. protectum over Heliomenina Gistel, 1848 nom. oblitum (Curculionidae), Naupactini Gistel, 1848 nom. protectum over Iphiini Schonherr, 1823 nom. oblitum (Curculionidae), Cleonini Schonherr, 1826 nom. protectum over Geomorini Schonherr, 1823 nom. oblitum (Curculionidae), Magdalidini Pascoe, 1870 nom. protectum over Scardamyctini Gistel, 1848 nom. oblitum (Curculionidae), Agrypninae/-ini Candeze, 1857 nom. protecta over Adelocerinae/-ini Gistel, 1848 nom. oblita and Pangaurinae/-ini Gistel, 1856 nom. oblita (Elateridae), Prosternini Gistel, 1856 nom. protectum over Diacanthini Gistel, 1848 nom. oblitum (Elateridae), Calopodinae Costa, 1852 nom. protectum over Sparedrinae Gistel, 1848 nom. oblitum (Oedemeridae), Adesmiini Lacordaire, 1859 nom. protectum over Macropodini Agassiz, 1846 nom. oblitum (Tenebrionidae), Bolitophagini Kirby, 1837 nom. protectum over Eledonini Billberg, 1820 nom. oblitum (Tenebrionidae), Throscidae Laporte, 1840 nom. protectum over Stereolidae Rafinesque, 1815 nom. oblitum (Throscidae) and Lophocaterini Crowson, 1964 over Lycoptini Casey, 1890 nom. oblitum (Trogossitidae); Monotoma Herbst, 1799 nom. protectum over Monotoma Panzer, 1792 nom. oblitum (Monotomidae); Pediacus Shuckard, 1839 nom. protectum over Biophloeus Dejean, 1835 nom. oblitum (Cucujidae), Pachypus Dejean, 1821 nom. protectum over Pachypus Billberg, 1820 nom. oblitum (Scarabaeidae), Sparrmannia Laporte, 1840 nom. protectum over Leocaeta Dejean, 1833 nom. oblitum and Cephalotrichia Hope, 1837 nom. oblitum (Scarabaeidae).

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Family-group names in Coleoptera (Insecta)

Summary

The phylogeny of carabid tribes is examined with sequences of 18S ribosomal DNA from eighty-four carabids representing forty-seven tribes, and fifteen outgroup taxa. Parsimony, distance and maximum likelihood methods are used to infer the phylogeny. Although many clades established with morphological evidence are present in all analyses, many of the basal relationships in carabids vary from analysis to analysis. These deeper relationships are also sensitive to variation in the sequence alignment under different alignment conditions. There is moderate evidence against the monophyly of Migadopini + Amarotypini, Scaritini + Clivinini, Bembidiini and Brachinini. Psydrini are not monophyletic, and consist of three distinct lineages (Psydrus, Laccocenus and a group of austral psydrines, from the Southern Hemisphere consisting of all the subtribes excluding Psydrina). The austral psydrines are related to Harpalinae plus Brachinini. The placements of many lineages, including Gehringia, Apotomus, Omophron, Psydrus and Cymbionotum, are unclear from these data. One unexpected placement, suggested with moderate support, is Loricera as the sister group to Amarotypus. Trechitae plus Patrobini form a monophyletic group. Brachinini probably form the sister group to Harpalinae, with the latter containing Pseudomorpha, Morion and Cnemalobus. The most surprising, well supported result is the placement of four lineages (Cicindelinae, Rhysodinae, Paussinae and Scaritini) as near relatives of Harpalinae + Brachinini. Because these four lineages all have divergent 18S rDNA, and thus have long basal branches, parametric bootstrapping was conducted to determine if their association and placement could be the result of long branch attraction. Simulations on model trees indicate that, although their observed association might be due to long branch attraction, there was no evidence that their placement near Harpalinae could be so explained. These simulations also suggest that 18S rDNA might not be sufficient to infer basal carabid relationships.

Phylogeny of carabid beetles as inferred from 18S ribosomal DNA (Coleoptera: Carabidae)

We consider the possibility of an extensive invasional meltdown occurring in central North America involving eleven Eurasian species. The scenario begins with the potential co-facilitation between the European earthworm Lumbricus terrestris and European buckthorn, Rhamnus cathartica. Once introduced, European buckthorn has served as the overwintering host for two important invasive crop pests, oat crown rust, Puccinea coronata and the soybean aphid, Aphis glycines. The spread of R. cathartica itself may have been aided by seed dispersal by the European starling, Sturnus vulgaris, and the presence of L. terrestris has likely facilitated the invasion of Bipalium adventitium, an Asian predatory flatworm that specializes on earthworms. Beyond this, the soybean aphid is consumed by a number of introduced species, including the lady beetle Harmonia axyridis, the ground beetle Agonum muelleri and the parasitoid Aphelinus certus. We hypothesize that the presence of soybean aphid increases regional abundances of these species. We discuss both the evidence for this multi-species invasional meltdown scenario and potential implications of meltdown dynamics for invasive species management. The particular management issues that we discuss are: (1) opportunities for managing multiple invasive species simultaneously by targeting facilitator species, and (2) implications of meltdown dynamics for biological control introductions against the soybean aphid.

European buckthorn and Asian soybean aphid as components of an extensive invasional meltdown in North America

Dealing with coleoptera, this is one of a series which presents New Zealand's invertebrate fauna. Each volume contains a checklist of the taxa, an introduction to the group, descriptions of the taxa, and illustrations.

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Coleoptera: family-group review and keys to identification

Effects of forest management practices on the diversity of ground-occurring beetles in mixed northern hardwood forests of the Great Lakes Region

All species-group names of Trachypachidae, Rhysodidae, and Carabidae (including cicindelincs) correctly recorded from America north of Mexico are catalogued with state and province records. Valid names are listed with the author(s), date of publication, and page citation in their current and original combinations while all synonyms are provided in their original combinations. Genus-group names are recorded with the author(s), date of publication, page citation, type species, and kind of type species fixation. Species groups were preferred to subgenera but subscneric names are also listed.The following nomenclatural changes are proposed and discussed: Bembidion neocoerulescens Bousquet, new replacement name for B. coerulescens Van Dyke, 1925; Chlaenius circumcinctus Say, 1830 for C. perplexus Dejean, 1831; Cyclotrachelus dejeanellus (Csiki, 1930) for C. morio (Dejean, 1828); Cyclotrachelus freitagi Bousquet, new replacement name for C. obsoletus (Say, 1830); Dyschirius aeneolus LeConte, 1850 for D. frigidus Mannerheim, 1853; Harpalus laevipes Zetterstedt, 1828 for H. quadripunctatus Dejean, 1829; Harpalus providens Casey, 1914 for H. viduus LeConte, 1865; Harpalus reversus Casey, 1924 for H. funerarius Csiki, 1932; Notiophilus sierranus Casey, 1920 for N. obscurus Fall, 1901; Pseudamara Lindroth, 1968 for Disamara Lindroth, 1976; Pterostichus trinarius (Casey, 1918) for P. ohionis Csiki, 1930; Stenolophus carbo Bousquet, new replacement name for S. carbonarius (Dejean, 1829).Thirty-six new synonyms are established and seven, considered as questionable, are confirmed. They are (with the valid names in parentheses): Agonothorax planipennis Motschulsky, 1850 (= ? Agonum affine Kirby, 1837); Platynus variolatus LeConte, 1851 (= Agonum limbatum Motschulsky, 1845); Agonum nitidum Harris, 1869 (= ? Agonum melanarium Dejean, 1828); Amerinus fuscicornis Casey, 1914 and A. longipennis Casey, 1914 (= Amerinus linearis (LeConte, 1863)); Apristus fuscipennis Motschulsky, 1864 (= Apristus latens LeConte, 1848); Batenus aeneolus Motschulsky, 1865 (= Agonum exaratum (Mannerheim, 1853)); Brachystylus curtipennis Motschulsky, 1859 (= Pterostichus congestus (Menetries, 1843)); Brachystylus parallelus Motschulsky, 1859 (= ? Pterostichus californicus (Dejean, 1828)); Cratacanthus cephalotes Casey, 1914, C. subovalis Casey, 1914, and C. texanus Casey, 1884 (= Cratacanthus dubius (Palisot de Beauvois, 1811)); Cymindis comma T.W. Harris, 1869 (= ? Cymindis limbatus Dejean, 1831); Feronia praetermissa Chaudoir, 1868 (= Pterostichus commutabilis (Motschulsky, 1866)); Galerita angusticeps Casey, 1920 (= Galerita janus (Fabricius, 1792)); Gonoderus cordicollis Motschulsky 1859 (= Pterostichus tristis (Dejean, 1828)); Anisodactylus alternans LeConte, 1851 (= Anisodactylus alternans (Motschulsky, 1845)); Hypherpes spissitarsis Casey, 1918 (= Pterostichus tarsalis LeConte, 1873); Lebia brunnicollis Motschulsky, 1864 (= Lebia lobulata LeConte, 1863); Lebia subfigurata Motschulsky, 1864 and L. sublimbata Motschulsky, 1864 (= Lebia analis Dejean, 1825); Lophoglossus bispiculatus Casey, 1913 and L. illini Casey, 1913 (= Lophoglossus scrutator (LeConte, 1848)); Platysma leconteianum Lutshnik, 1922 (= Pterostichus commutabilis (Motschulsky, 1866)); Loxandrus iris Motschulsky, 1866(= Loxandrus rectus (Say, 1823)); Masoreus americanus Motschulsky, 1864 (= Stenolophus rotundicollis (Haldeman, 1843)); Notaphus laterimaculatus Motschulsky, 1859 (= Bembidion approximatum (LeConte, 1852)); Notiophilus cribrilaterus Motschulsky, 1864 (= Notiophilus novemstriatus LeConte, 1848); Omaseus brevibasis Casey, 1924 (= Pterostichus luctuosus (Dejean, 1828)); Notaphus incertus Motschulsky, 1845 (= Bembidion breve (Motschulsky, 1845)); Peryphus concolor Motschulsky, 1850 (= Bembidion platynoides Hayward, 1897); Peryphus erosus Motschulsky, 1850 (= Bembidion transversale Dejean, 1831); Peryphus subinflatus Motschulsky, 1859 (= Bembidion petrosum petrosum Gebler, 1833); Planesus fuscicollis Motschulsky, 1865 and P. laevigatas Motschulsky, 1865 (= Cymindis platicollis (Say, 1823)); Poecilus pimalis Casey, 1913 (= Poecilus diplophryus Chaudoir, 1876); Pterostichus arizonicus Schaeffer, 1910 (= Ophryogaster flohri Bates, 1882); Pterostichus sequoiarum Casey, 1913 (= Pterostichus tarsalis LeConte, 1873); Scaphinotus grandis Gistel, 1857 (= ? Scaphinotus unicolor unicolor (Fabricius, 1787)); Stenocrepis chalcas Bates, 1882 and S. chalcochrous Chaudoir, 1883 (= Stenocrepis texana (LeConte, 1863)); Stenolophus humeralis Motschulsky, 1864 (= Stenolophus plebejus Dejean, 1829); and Stenolophus laticollis Motschulsky, 1864 (= Stenolophus ochropezus (Say, 1823)).Olisthopus iterans Casey, 1913 and Pterostichus illustris LeConte, 1851, listed as junior synonyms of O. parmatus (Say, 1823) and P. congestus (Menetries, 1843), respectively, are considered in the present work as valid species.The type species (listed in parentheses) of the following 14 genus-group taxa are designated for the first time: Circinalidia Casey, 1920 (Agonum aeruginosum Dejean, 1828); Evolenes LeConte, 1853 (Oodes exaratus Dejean, 1831); Leucagonum Casey, 1920 (Agonum maculicolle Dejean, 1828); Megaliridia Casey, 1920 (Cychrus viduus Dejean, 1826); Megalostylus Chaudoir, 1843 (Feronia lucidula Dejean, 1828 = Feronia recta Say, 1823); Micragra Chaudoir, 1872 (Micragra lissonota Chaudoir, 1872); Onota Chaudoir, 1872 (Onota bicolor Chaudoir, 1872); Oodiellus Chaudoir, 1882 (Oodiellus mexicanus Chaudoir, 1882 = Anatrichis alutacea Bates, 1882); Oxydrepanus Putzeys, 1866 (Dyschirius rufus Putzeys, 1846); Paranchomenus Casey, 1920 (Platynus stygicus LeConte, 1854 = Anchomenus mannerheimii Dejean, 1828); Pemphus Motschulsky, 1866 (Cychrus velutinus Menetries, 1843); Peronoscelis Chaudoir, 1872 (Tetragonoderus figuratus Dejean, 1831); Rhombodera Reiche, 1842 (Rhombodera virgata Reiche, 1842 = Lebia trivittata Dejean, 1831); and Stenous Chaudoir, 1857 (Oodes cupreus Chaudoir, 1843).Two new family-group names are proposed, Cnemalobini (= Cnemacanthini of authors) based on Cnemalobus Guerin-Meneville, 1839 and Loxandrini based on Loxandrus LeConte, 1852.The work also includes a synopsis of all extant world carabid tribes, a bibliography of all original descriptions, a full taxonomic index, and, as appendices, lists of nomina nuda and unjustified emendations, and annotated lists of species incorrectly or doubtfully recorded from America north of Mexico and of new North American records.

Y. Bousquet

Papers

Catalogue of the Geadephaga (Coleoptera: Trachypachidae, Rhysodidae, Carabidae including Cicindelini) of America north of Mexico