Preface to the Princeton Landmarks in Biology Edition vii Preface xi Symbols Used xiii 1. The Importance of Islands 3 2. Area and Number of Speicies 8 3. Further Explanations of the Area-Diversity Pattern 19 4. The Strategy of Colonization 68 5. Invasibility and the Variable Niche 94 6. Stepping Stones and Biotic Exchange 123 7. Evolutionary Changes Following Colonization 145 8. Prospect 181 Glossary 185 References 193 Index 201

The Theory of Island Biogeography

In 1997, the global value of ecosystem services was estimated to average $33 trillion/yr in 1995 $US ($46 trillion/yr in 2007 $US). In this paper, we provide an updated estimate based on updated unit ecosystem service values and land use change estimates between 1997 and 2011. We also address some of the critiques of the 1997 paper. Using the same methods as in the 1997 paper but with updated data, the estimate for the total global ecosystem services in 2011 is $125 trillion/yr (assuming updated unit values and changes to biome areas) and $145 trillion/yr (assuming only unit values changed), both in 2007 $US. From this we estimated the loss of eco-services from 1997 to 2011 due to land use change at $4.3–20.2 trillion/yr, depending on which unit values are used. Global estimates expressed in monetary accounting units, such as this, are useful to highlight the magnitude of eco-services, but have no specific decision-making context. However, the underlying data and models can be applied at multiple scales to assess changes resulting from various scenarios and policies. We emphasize that valuation of eco-services (in whatever units) is not the same as commodification or privatization. Many eco-services are best considered public goods or common pool resources, so conventional markets are often not the best institutional frameworks to manage them. However, these services must be (and are being) valued, and we need new, common asset institutions to better take these values into account.

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Changes in the global value of ecosystem services

Mapping ecosystem service supply, demand and budgets

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Input Output Analysis Foundations And Extensions

In order to use environmental models effectively for management and decision-making, it is vital to establish an appropriate level of confidence in their performance. This paper reviews techniques available across various fields for characterising the performance of environmental models with focus on numerical, graphical and qualitative methods. General classes of direct value comparison, coupling real and modelled values, preserving data patterns, indirect metrics based on parameter values, and data transformations are discussed. In practice environmental modelling requires the use and implementation of workflows that combine several methods, tailored to the model purpose and dependent upon the data and information available. A five-step procedure for performance evaluation of models is suggested, with the key elements including: (i) (re)assessment of the model's aim, scale and scope; (ii) characterisation of the data for calibration and testing; (iii) visual and other analysis to detect under- or non-modelled behaviour and to gain an overview of overall performance; (iv) selection of basic performance criteria; and (v) consideration of more advanced methods to handle problems such as systematic divergence between modelled and observed values.

Position paper: Characterising performance of environmental models

Landscapes differ in their capacities to provide ecosystem goods and services, which are the benefits humans obtain from nature. Structures and functions of ecosystems needed to sustain the provision of ecosystem services are altered by various human activities. In this paper, a concept for the assessment of multiple ecosystem services is proposed as a basis for discussion and further development of a respective evaluation instrument. Using quantitative and qualitative assessment data in combination with land cover and land use information originated from remote sensing and GIS, impacts of human activities can be evaluated. The results reveal typical patterns of different ecosystems‘ capacities to provide ecosystem services. The proposed approach thus delivers useful integrative information for environmental management and landscape planning, aiming at a sustainable use of services provided by nature. The research concept and methodological framework presented here for discussion have initially been applied in different case studies and shall be developed further to provide a useful tool for the quantification and spatial modelling of multiple ecosystem services in different landscapes. An exemplary application of the approach dealing with food provision in the Halle-Leipzig region in Germany is presented. It shows typical patterns of ecosystem service distribution around urban areas. As the approach is new and still rather general, there is great potential for improvement, especially with regard to a data-based quantification of the numerous hypotheses, which were formulated as base for the assessment. Moreover, the integration of more detailed landscape information on different scales will be needed in future in order to take the heterogeneous distribution of landscape properties and values into account. Therefore, the purpose of this paper is to foster critical discussions on the methodological development presented here.

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Landscapes' Capacities to Provide Ecosystem Services - a Concept for Land-Cover Based Assessments

The high variety of ecosystem service categorisation systems, assessment frameworks, indicators, quantification methods and spatial localisation approaches allows scientists and decision makers to harness experience, data, methods and tools. On the other hand, this variety of concepts and disagreements among scientists hamper an integration of ecosystem services into contemporary environmental management and decision making. In this article, the current state of the art of ecosystem service science regarding spatial localisation, indication and quantification of multiple ecosystem service supply and demand is reviewed and discussed. Concepts and tables for regulating, provisioning and cultural ecosystem service definitions, distinguishing between ecosystem service potential supply (stocks), flows (real supply) and demands as well as related indicators for quantification are provided. Furthermore, spatial concepts of service providing units, benefitting areas, spatial relations, rivalry, spatial and temporal scales are elaborated. Finally, matrices linking CORINE land cover types to ecosystem service potentials, flows, demands and budget estimates are provided. The matrices show that ecosystem service potentials of landscapes differ from flows, especially for provisioning ecosystem services.

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Felix Müller

Papers

Mapping ecosystem service supply, demand and budgets

Landscapes' Capacities to Provide Ecosystem Services - a Concept for Land-Cover Based Assessments

Ecosystem service potentials, flows and demands-concepts for spatial localisation, indication and quantification

Rural-urban gradient analysis of ecosystem services supply and demand dynamics

Interactions of ecosystem properties, ecosystem integrity and ecosystem service indicators—A theoretical matrix exercise