Geological atlas of western and central Europe

A correlation of tectonic units of the Alpine-Carpathian-Dinaridic system of orogens, including the substrate of the Pannonian and Transylvanian basins, is presented in the form of a map. Combined with a series of crustal-scale cross sections this correlation of tectonic units yields a clearer picture of the three-dimensional architecture of this system of orogens that owes its considerable complexity to multiple overprinting of earlier by younger deformations.

/pdf/the-alpine-carpathian-dinaridic-orogenic-system-correlation-2gm3qeefzn.pdf

The Alpine-Carpathian-Dinaridic orogenic system: correlation and evolution of tectonic units

Reconstructing the origin and evolution of land plants and their algal relatives is a fundamental problem in plant phylogenetics, and is essential for understanding how critical adaptations arose, including the embryo, vascular tissue, seeds, and flowers. Despite advances in molecular systematics, some hypotheses of relationships remain weakly resolved. Inferring deep phylogenies with bouts of rapid diversification can be problematic; however, genome-scale data should significantly increase the number of informative characters for analyses. Recent phylogenomic reconstructions focused on the major divergences of plants have resulted in promising but inconsistent results. One limitation is sparse taxon sampling, likely resulting from the difficulty and cost of data generation. To address this limitation, transcriptome data for 92 streptophyte taxa were generated and analyzed along with 11 published plant genome sequences. Phylogenetic reconstructions were conducted using up to 852 nuclear genes and 1,701,170 aligned sites. Sixty-nine analyses were performed to test the robustness of phylogenetic inferences to permutations of the data matrix or to phylogenetic method, including supermatrix, supertree, and coalescent-based approaches, maximum-likelihood and Bayesian methods, partitioned and unpartitioned analyses, and amino acid versus DNA alignments. Among other results, we find robust support for a sister-group relationship between land plants and one group of streptophyte green algae, the Zygnematophyceae. Strong and robust support for a clade comprising liverworts and mosses is inconsistent with a widely accepted view of early land plant evolution, and suggests that phylogenetic hypotheses used to understand the evolution of fundamental plant traits should be reevaluated.

/pdf/phylotranscriptomic-analysis-of-the-origin-and-early-4cs38v80rr.pdf

Phylotranscriptomic analysis of the origin and early diversification of land plants

The green lineage (Viridiplantae) comprises the green algae and their descendants the land plants, and is one of the major groups of oxygenic photosynthetic eukaryotes. Current hypotheses posit the early divergence of two discrete clades from an ancestral green flagellate. One clade, the Chlorophyta, comprises the early diverging prasinophytes, which gave rise to the core chlorophytes. The other clade, the Streptophyta, includes the charophyte green algae from which the land plants evolved. Multi-marker and genome scale phylogenetic studies have greatly improved our understanding of broad-scale relationships of the green lineage, yet many questions persist, including the branching orders of the prasinophyte lineages, the relationships among core chlorophyte clades (Chlorodendrophyceae, Ulvophyceae, Trebouxiophyceae and Chlorophyceae), and the relationships among the streptophytes. Current phylogenetic hypotheses provide an evolutionary framework for molecular evolutionary studies and comparative genomics. This review summarizes our current understanding of organelle genome evolution in the green algae, genomic insights into the ecology of oceanic picoplanktonic prasinophytes, molecular mechanisms underlying the evolution of complexity in volvocine green algae, and the evolution of genetic codes and the translational apparatus in green seaweeds. Finally, we discuss molecular evolution in the streptophyte lineage, emphasizing the genetic facilitation of land plant origins.

/pdf/phylogeny-and-molecular-evolution-of-the-green-algae-4244e09mtu.pdf

Phylogeny and Molecular Evolution of the Green Algae

Establishing the timescale of early land plant evolution is essential for testing hypotheses on the coevolution of land plants and Earth's System. The sparseness of early land plant megafossils and stratigraphic controls on their distribution make the fossil record an unreliable guide, leaving only the molecular clock. However, the application of molecular clock methodology is challenged by the current impasse in attempts to resolve the evolutionary relationships among the living bryophytes and tracheophytes. Here, we establish a timescale for early land plant evolution that integrates over topological uncertainty by exploring the impact of competing hypotheses on bryophyte-tracheophyte relationships, among other variables, on divergence time estimation. We codify 37 fossil calibrations for Viridiplantae following best practice. We apply these calibrations in a Bayesian relaxed molecular clock analysis of a phylogenomic dataset encompassing the diversity of Embryophyta and their relatives within Viridiplantae. Topology and dataset sizes have little impact on age estimates, with greater differences among alternative clock models and calibration strategies. For all analyses, a Cambrian origin of Embryophyta is recovered with highest probability. The estimated ages for crown tracheophytes range from Late Ordovician to late Silurian. This timescale implies an early establishment of terrestrial ecosystems by land plants that is in close accord with recent estimates for the origin of terrestrial animal lineages. Biogeochemical models that are constrained by the fossil record of early land plants, or attempt to explain their impact, must consider the implications of a much earlier, middle Cambrian-Early Ordovician, origin.

/pdf/the-timescale-of-early-land-plant-evolution-2e9rvqrjsv.pdf

The timescale of early land plant evolution

Application of the topographic position index to heterogeneous landscapes

Colonization of the land by plants most likely occurred in a stepwise fashion starting in the Mid-Ordovician. The earliest flora of bryophyte-like plants appears to have been cosmopolitan and dominated the planet, relatively unchanged, for some 30 million years. It is represented by fossilized dispersed cryptospores and fragmentary plant remains. In the Early Silurian, cryptospore abundance and diversity diminished abruptly as trilete spores appeared, became abundant, and underwent rapid diversification. This change coincides approximately with the appearance of vascular plant megafossils and probably represents the origin and adaptive radiation of vascular plants. We have obtained a diverse trilete spore occurrence from the Late Ordovician that suggests that vascular plants originated and diversified earlier than previously hypothesized, in Gondwana, before migrating elsewhere and secondarily diversifying.

/pdf/origin-and-radiation-of-the-earliest-vascular-land-plants-nzveov99hf.pdf

Origin and Radiation of the Earliest Vascular Land Plants

Abstract Multidisciplinary studies undertaken within the EU-funded PACE Network have permitted a new 3-D reassessment of the relationships between the principal crustal blocks abutting Baltica along the Trans-European Suture Zone (TESZ). The simplest model indicates that accretion was in three stages: end-Cambrian accretion of the Bruno-Silesian, Łysogóry and Małopolska terranes; late Ordovician accretion of Avalonia, and early Carboniferous accretion of the Armorican Terrane Assemblage (ATA), which had coalesced during Late Devonian — Early Carboniferous time. All these accreted blocks contain similar Neoproterozoic basement indicating a peri-Gondwanan origin: Palaeozoic plume-influenced metabasite geochemistry in the Bohemian Massif in turn may explain their progressive separation from Gondwana before their accretion to Baltica, although separation of the Bruno-Silesian and related blocks from Baltica during the Cambrian is contentious. Inherited ages from both the Bruno-Silesian crustal block and Avalonia contain a 1.5 Ga ‘Rondonian’ component arguing for proximity to the Amazonian craton at the end of the Neoproterozoic: such a component is absent from Armorican terranes, which suggests that they have closer affinities with the West African craton. Models showing the former locations of these terranes and the larger continents from which they rifted, or to which they became attached, must conform to the above constraints, as well as those provided by palaeomagnetic data. Hence, at the end of the Proterozoic and in the early Palaeozoic, these smaller terranes, some of which contain Neoproterozoic ophiolitic marginal basin and magmatic arc remnants, probably occurred within the end-Proterozoic supercontinent as part of a ‘Pacific-type’ margin, which became dismembered and relocated as the supercontinent fragmented.

/pdf/palaeozoic-amalgamation-of-central-europe-an-introduction-232nc65g92.pdf

Palaeozoic amalgamation of Central Europe: an introduction and synthesis of new results from recent geological and geophysical investigations

A global Chitinozoan biozonation for the Silurian Period is proposed. Each biozone is an interval range defined by the first occurrence of an index species, selected from well-defined species with a relatively short biostratigraphic range. To be included in the scheme, index species must have been recorded from at least two major Silurian palaeocontinents where usable Chitinozoa assemblages have been studied, that is, Avalonia—Baltica (which had already undergone collision by the Silurian), Laurentia, Gondwana and Southern China. Seventeen biozones are identified with seven in the Llandovery: the fragilis, postrobusta, electa, maennili, alargada, dolioliformis and longicollis biozones; four in the Wenlock: the margaritana, cingulata, pachycephala and lycoperdoides biozones; three in the Ludlow: the elongata, philipi and barrandei biozones; and three in the Přidoli: the kosovensis, elegans and superba biozones. Chronostratigraphic calibration is partly provided by reference to the range of the appropriate Chitinozoa index species in the global stratotype sections and points (GSSP) of the Silurian series, e.g. in Bohemia (Czech Republic) for the Přidoli and the Welsh Borderland in England for the Ludlow and part of the Wenlock. When this information was not available, independent biostratigraphic control was provided by calibration with graptolite biozones or in a few cases, conodont or trilobite biozonal schemes. The index and most characteristic species of each biozone are illustrated and their total stratigraphie range provided.

A global Chitinozoa biozonation for the Silurian

Palaeozoic Amalgamation of Central Europe summarizes recent research designed to clarify the timing, geometry and processes by which discrete terranes of Central Europe became amalgamated during the Palaeozoic Era. The area studied extends from the southern North Sea to Central Poland along the Trans-European Suture Zone, covering much of Germany, Denmark, Belgium, the Czech Republic and Poland.

The 16 papers within the volume are divided into five sections: biostratigraphic/provenance evidence; isotopic constraints; petrological and geochemical evidence; structural evolution; seismic traverses and deep crustal structure. The first section contains papers summarizing continent-specific micropalaeontological and sediment provenance information backing current debates about microcontinent derivation and timing of their accretions to the proto-European continent, Baltica. The section on isotopic constraints discusses the use of isotopic dating to constrain the timing of accretions of rock units exposed in the northern Bohemian Massif, while the following section has more detailed studies of metamorphosed ophiolitic complexes adjoining palaeosutures in the same area. The two papers on the structural evolution of the area contrast a detailed review of the structural evolution of the Sudetes, with a broader, more regionally based hypothesis for the structural evolution of all Central Europe. The final section discusses models based on extensive seismic traverses in contrasting parts of the area - Belgium, the southern North Sea and Poland. This wide-ranging study thus encapsulates the most up-to-date ideas on the Palaeozoic amalgamation of Central Europe from the leading international researchers in the field.

The volume will be of interest to those earth scientists in industry and academia with a broad-based interest in the construction of the European continent, primarily biostratigraphers, geophysicists, structural geologists and geochemists.

http://www.gbv.de/dms/goettingen/360810322.pdf

Jacques Verniers

Papers

Application of the topographic position index to heterogeneous landscapes

Origin and Radiation of the Earliest Vascular Land Plants

Palaeozoic amalgamation of Central Europe: an introduction and synthesis of new results from recent geological and geophysical investigations

A global Chitinozoa biozonation for the Silurian

Palaeozoic Amalgamation of Central Europe