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The influence of diet on the distribution of nitrogen isotopes in animals was investigated by analyzing animals grown in the laboratory on diets of constant nitrogen isotopic composition. 
The isotopic composition of the nitrogen in an animal reflects the nitrogen isotopic composition of its diet. The δ^(15)N values of the whole bodies of animals are usually more positive than those of their diets. Different individuals of a species raised on the same diet can have significantly different δ^(15)N values. The variability of the relationship between the δ^(15)N values of animals and their diets is greater for different species raised on the same diet than for the same species raised on different diets. Different tissues of mice are also enriched in ^(15)N relative to the diet, with the difference between the δ^(15)N values of a tissue and the diet depending on both the kind of tissue and the diet involved. The δ^(15)N values of collagen and chitin, biochemical components that are often preserved in fossil animal remains, are also related to the δ^(15)N value of the diet. 
The dependence of the δ^(15)N values of whole animals and their tissues and biochemical components on the δ^(15)N value of diet indicates that the isotopic composition of animal nitrogen can be used to obtain information about an animal's diet if its potential food sources had different δ^(15)N values. The nitrogen isotopic method of dietary analysis probably can be used to estimate the relative use of legumes vs non-legumes or of aquatic vs terrestrial organisms as food sources for extant and fossil animals. However, the method probably will not be applicable in those modern ecosystems in which the use of chemical fertilizers has influenced the distribution of nitrogen isotopes in food sources. 
The isotopic method of dietary analysis was used to reconstruct changes in the diet of the human population that occupied the Tehuacan Valley of Mexico over a 7000 yr span. Variations in the δ^(15)C and δ^(15)N values of bone collagen suggest that C_4 and/or CAM plants (presumably mostly corn) and legumes (presumably mostly beans) were introduced into the diet much earlier than suggested by conventional archaeological analysis.

Influence of Diet On the Distribtion of Nitrogen Isotopes in Animals

A History of Archaeological Thought.

Bioarchaeology: Interpreting behavior from the human skeleton

The human remains

The effects of Egyptian imperial expansion into Nubia during the New Kingdom Period (1,550–1,069 BC) have been debated. Here, the impacts of the Egyptian Empire are investigated through an examination of osteological indicators of activity at the archaeological site of Tombos. Entheseal changes to fibrocartilaginous attachment sites and osteoarthritis are examined to infer what types of physical activities this colonial town was engaging in. Many of the skeletal remains at Tombos were comingled due to looting in antiquity; undisturbed burials are presented as a subsample of the population (n = 28) in which age, sex, and body size can be considered. The total sample (n = 85) is then analyzed to better understand overall levels of activity. A number of Nile River Valley bioarchaeological samples are used as points of comparison to the Tombos population. Results indicate that the inhabitants of Tombos had relatively low entheseal remodeling scores; this is highlighted when Tombos is juxtaposed with comparative samples, particularly in men. Furthermore, osteoarthritis, as assessed by eburnation, was also markedly infrequent at Tombos. Collectively, these results indicate a relatively low level of activity and support the hypothesis that Tombos may have served as an administrative center. Am J Phys Anthropol, 2012. © 2012 Wiley Periodicals, Inc.

Activity patterns in New Kingdom Nubia: An examination of entheseal remodeling and osteoarthritis at Tombos

Located 10 km south of the Third Cata- ract of the Nile River, the ancient city of Kerma was once capital to the second largest state in Africa. The Eastern Cemetery at Kerma (4 km east of city center) encompasses 801 hectares and was used over a period of 1,500 years (3,200-1,500 BC). Excavated in the early 20th century by George Reisner, the cemetery contained an estimated 20,000-40,000 individuals. Reisner classi- fied these burials into multiple categories, including chiefs and human sacrifices, based on burial position and grave goods. This study investigates the skeletal embodiment of social inequality by examining variation in entheseal severity between the Kerma burial classifications. Seventeen entheses were examined using the Hawkey and Merbs (1995) scoring method (n 5 205 individuals); age, sex, and body size variables were con- sidered by employing Mann-Whitney U tests and partial Spearman's correlations. This analysis suggests that sig- nificant differences in entheseal changes existed between select burial types. Specifically, "corridor sac- rifices" had significantly higher rates of entheseal changes while "chiefs" and "subsidiary burials" had simi- lar entheseal changes; furthermore, within these burial categories, males had higher entheseal scores despite body size controls. The elevated entheseal changes in the sacrificial burials may be due to an intensive agro- pastoral lifestyle or other demanding forms of manual labor. In conclusion, the disparity of entheseal markers between burial subgroups at Kerma might reflect a degree of social inequality within this state level society. This bioarchaeological research informs our understand- ing of socially-defined categories of persons as well as everyday life in Ancient Kerma. Am J Phys Anthropol

Elucidating inequality in Nubia: An examination of entheseal changes at Kerma (Sudan)

Isotopic Approaches to Mobility in Northern Africa: A Bioarchaeological Examination of Egyptian/Nubian Interaction in the Nile Valley

Bioarchaeological perspectives of collapse scenarios have contributed to our understanding of the persistence and regeneration of communities after the fall of states and empires. We suggest that the osteological examination of everyday lived experience can further elucidate local lifeways during these transitory periods. Focusing on ancient Nubia after the decline of the colonial Egyptian New Kingdom Empire, we examine musculoskeletal activity patterns (entheseal changes) and traumatic injury in remains uncovered at Tombos, a cemetery at the Third Cataract of the Nile. We find evidence that entheseal changes increased at Tombos from the New Kingdom Period to the postcolonial Third Intermediate/Napatan Periods; however, accidental trauma does not significantly increase between the two periods. With regard to the elevated entheseal changes, we suggest that Nubians may have been engaged in quarrying, agropastoralism, and construction efforts. Furthermore, there is evidence to suggest that this population was biologically resilient, which may have limited accidental/occupational traumatic injury. We view these changes to the Tombos quotidian experience from an agent-centered perspective; we posit that during the postcolonial period, Nubians revitalized their sociopolitical and economic systems through these everyday practices, which contributed to the independent and powerful Napatan state.

Everyday Life after Collapse: a Bioarchaeological Examination of Entheseal Change and Accidental Injury in Postcolonial Nubia

Sex estimation is an important part of creating a biological profile for skeletal remains in forensics. The commonly used methods for developing sex estimation equations are discriminant function analysis (DFA) and logistic regression (LogR). LogR equations provide a probability of the predicted sex, while DFA relies on cutoff points to segregate males and females, resulting in a rigid dichotomization of the sexes. This is problematic because sexual dimorphism exists along a continuum and there can be considerable overlap in trait expression between the sexes. In this study, we used humeral measurements to compare the performance of DFA and LogR and found them to be very similar under multiple conditions. The overall cross-validated (leave-one-out) accuracy of DFA (75.76-95.14%) was slightly higher than LogR (75.76-93.82%) for simple and multiple variable equations, and also performed better under varying sample sizes (94.03% vs. 93.78%). Three of five DFA equations outperformed LogR under the B index, while all five LogR equations outperformed the DFA equations under the Q index. Both methods saw an improvement in overall accuracy (DFA: 86.74-95.79%; LogR: 86.74-95.76%) when individuals with a classification probability lower than 0.80 were excluded. Additionally, we propose a method for calculating additional cutoff points (PMarks) based on posterior probability values. In conclusion, we recommend using LogR over DFA due to the increased flexibility, robusticity, and benefits for future users of the statistical models; however, if DFA is preferred, use of the proposed PMarks facilitates future analysis while avoiding unnecessary dichotomization.

https://onlinelibrary.wiley.com/doi/pdf/10.1111/1556-4029.14482

Sarah A. Schrader

Papers

Activity patterns in New Kingdom Nubia: An examination of entheseal remodeling and osteoarthritis at Tombos

Elucidating inequality in Nubia: An examination of entheseal changes at Kerma (Sudan)

Isotopic Approaches to Mobility in Northern Africa: A Bioarchaeological Examination of Egyptian/Nubian Interaction in the Nile Valley

Everyday Life after Collapse: a Bioarchaeological Examination of Entheseal Change and Accidental Injury in Postcolonial Nubia

Getting Rid of Dichotomous Sex Estimations: Why Logistic Regression Should be Preferred Over Discriminant Function Analysis.