Preface to the Princeton Landmarks in Biology Edition vii Preface xi Symbols Used xiii 1. The Importance of Islands 3 2. Area and Number of Speicies 8 3. Further Explanations of the Area-Diversity Pattern 19 4. The Strategy of Colonization 68 5. Invasibility and the Variable Niche 94 6. Stepping Stones and Biotic Exchange 123 7. Evolutionary Changes Following Colonization 145 8. Prospect 181 Glossary 185 References 193 Index 201

The Theory of Island Biogeography

Glacial melting in the Tibetan Plateau affects the water resources of millions of people. This study finds that—partly owing to changes in atmospheric circulations and precipitation patterns—the most intensive glacier shrinkage is in the Himalayan region, whereas glacial retreat in the Pamir Plateau region is less apparent.

Different glacier status with atmospheric circulations in Tibetan Plateau and surroundings

Prefaces Acknowledgements 1. Mountains and their climatological study 2. Geographical controls of mountain meteorological elements 3. Circulation systems related to orography 4. Climatic characteristics of mountains 5. Regional case studies 6. Mountain bioclimatology 7. Changes in mountain climates Appendix General index Author index.

Mountain Weather and Climate

Tree Rings And Climate

The Greater Himalayas hold the largest mass of ice outside polar regions and are the source of the 10 largest rivers in Asia. Rapid reduction in the volume of Himalayan glaciers due to climate change is occurring. The cascading effects of rising temperatures and loss of ice and snow in the region are affecting, for example, water availability (amounts, seasonality), biodiversity (endemic species, predator-prey relations), ecosystem boundary shifts (tree-line movements, high-elevation ecosystem changes), and global feedbacks (monsoonal shifts, loss of soil carbon). Climate change will also have environmental and social impacts that will likely increase uncertainty in water supplies and agricultural production for human populations across Asia. A common understanding of climate change needs to be developed through regional and local-scale research so that mitigation and adaptation strategies can be identified and implemented. The challenges brought about by climate change in the Greater Himalayas can only be addressed through increased regional collaboration in scientific research and policy making.

The melting Himalayas: cascading effects of climate change on water, biodiversity, and livelihoods.

As a mountainous country, Nepal is most susceptible to precipitation extremes and related hazards, including severe floods, landslides and droughts that cause huge losses of life and property, impact the Himalayan environment, and hinder the socioeconomic development of the country. Given that the countrywide assessment of such extremes is still lacking, we present a comprehensive picture of prevailing precipitation extremes observed across Nepal. First, we present the spatial distribution of daily extreme precipitation indices as defined by the Expert Team on Climate Change Detection, Monitoring and Indices (ETCCDMI) from 210 stations over the period of 1981–2010. Then, we analyze the temporal changes in the computed extremes from 76 stations, featuring long-term continuous records for the period of 1970–2012, by applying a non-parametric Mann−Kendall test to identify the existence of a trend and Sen’s slope method to calculate the true magnitude of this trend. Further, the local trends in precipitation extremes have been tested for their field significance over the distinct physio-geographical regions of Nepal, such as the lowlands, middle mountains and hills and high mountains in the west (WL, WM and WH, respectively), and likewise, in central (CL, CM and CH) and eastern (EL, EM and EH) Nepal. Our results suggest that the spatial patterns of high-intensity precipitation extremes are quite different to that of annual or monsoonal precipitation. Lowlands (Terai and Siwaliks) that feature relatively low precipitation and less wet days (rainy days) are exposed to high-intensity precipitation extremes. Our trend analysis suggests that the pre-monsoonal precipitation is significantly increasing over the lowlands and CH, while monsoonal precipitation is increasing in WM and CH and decreasing in CM, CL and EL. On the other hand, post-monsoonal precipitation is significantly decreasing across all of Nepal while winter precipitation is decreasing only over the WM region. Both high-intensity precipitation extremes and annual precipitation trends feature east−west contrast, suggesting significant increase over the WM and CH region but decrease over the EM and CM regions. Further, a significant positive trend in the number of consecutive dry days but significant negative trend in the number of wet (rainy) days are observed over the whole of Nepal, implying the prolongation of the dry spell across the country. Overall, the intensification of different precipitation indices over distinct parts of the country indicates region-specific risks of floods, landslides and droughts. The presented findings, in combination with population and environmental pressures, can support in devising the adequate region-specific adaptation strategies for different sectors and in improving the livelihood of the rural communities in Nepal.

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Rising Precipitation Extremes across Nepal

Based on comprehensive evaluations and analyses of existing literature and data sources, a review of geographical and ecological aspects of the upper timberline in the Himalayan mountain system is presented. Upper timberline elevations increase along two gradients: a NW-SE gradient corresponds to higher temperature sums at same elevations along the mountain arc. However, mean temperatures of the warmest month are higher at timberlines in the NW, which develop at lower elevations than expected since extreme winter cold, later snow melt, and shorter growing seasons overcompensate the advantage of higher summer temperatures. A second gradient is developed in peripheral-central direction from the Himalayan south slope to the Great Himalayan range and the Tibetan highlands. Increasing timberline elevations along this gradient are related to the combined effects of continentality and mass-elevation, both leading to higher temperature sums. Juniperus tibetica stands in S-Central Tibet even reach 4600–4800 m, the most elevated timberline in the northern hemisphere.

The Upper Timberline in the Himalayas, Hindu Kush and Karakorum: a Review of Geographical and Ecological Aspects

. Climate warming is expected to induce treelines to advance to higher elevations. Empirical studies in diverse mountain ranges, however, give evidence of both advancing alpine treelines and rather insignificant responses. The inconsistency of findings suggests distinct differences in the sensitivity of global treelines to recent climate change. It is still unclear where Himalayan treeline ecotones are located along the response gradient from rapid dynamics to apparently complete inertia. This paper reviews the current state of knowledge regarding sensitivity and response of Himalayan treelines to climate warming, based on extensive field observations, published results in the widely scattered literature, and novel data from ongoing research of the present authors. Several sensitivity indicators such as treeline type, treeline form, seed-based regeneration, and growth patterns are evaluated. Since most Himalayan treelines are anthropogenically depressed, observed advances are largely the result of land use change. Near-natural treelines are usually krummholz treelines, which are relatively unresponsive to climate change. Nevertheless, intense recruitment of treeline trees suggests a great potential for future treeline advance. Competitive abilities of seedlings within krummholz thickets and dwarf scrub heaths will be a major source of variation in treeline dynamics. Tree growth–climate relationships show mature treeline trees to be responsive to temperature change, in particular in winter and pre-monsoon seasons. High pre-monsoon temperature trends will most likely drive tree growth performance in the western and central Himalaya. Ecological niche modelling suggests that bioclimatic conditions for a range expansion of treeline trees will be created during coming decades.

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Do Himalayan treelines respond to recent climate change? An evaluation of sensitivity indicators

Little is known about how soil properties control tree growth at its upper limit. This paper reviews the state of knowledge and discusses the results specifically related to ecozones, to the scale-...

Udo Schickhoff

Papers

Rising Precipitation Extremes across Nepal

The Upper Timberline in the Himalayas, Hindu Kush and Karakorum: a Review of Geographical and Ecological Aspects

Do Himalayan treelines respond to recent climate change? An evaluation of sensitivity indicators

How do soil properties affect alpine treelines? : General principles in a global perspective and novel findings from Rolwaling Himal, Nepal

Himalayan treeline soil and foliar C:N:P stoichiometry indicate nutrient shortage with elevation