Connectivity, or the exchange of individuals among marine populations, is a central topic in marine ecology. For most benthic marine species with complex life cycles, this exchange occurs primarily during the pelagic larval stage. The small size of larvae coupled with the vast and complex fluid environment they occupy hamper our ability to quantify dispersal and connectivity. Evidence from direct and indirect approaches using geochemical and genetic techniques suggests that populations range from fully open to fully closed. Understanding the biophysical processes that contribute to observed dispersal patterns requires integrated interdisciplinary approaches that incorporate high-resolution biophysical modeling and empirical data. Further, differential postsettlement survival of larvae may add complexity to measurements of connectivity. The degree to which populations self recruit or receive subsidy from other populations has consequences for a number of fundamental ecological processes that affect population regulation and persistence. Finally, a full understanding of population connectivity has important applications for management and conservation.

Larval Dispersal and Marine Population Connectivity

The ocean circulation is a cause and consequence of fluid scale interactions ranging from millimeters to more than 10,000 km. Although the wind field produces a large energy input to the ocean, all but approximately 10% appears to be dissipated within about 100 m of the sea surface, rendering observations of the energy divergence necessary to maintain the full water-column flow difficult. Attention thus shifts to the physically different kinetic energy (KE) reservoirs of the circulation and their maintenance, dissipation, and possible influence on the very small scales representing irreversible molecular mixing. Oceanic KE is dominated by the geostrophic eddy field, and depending on the vertical structure (barotropic versus low-mode baroclinic), direct and inverse energy cascades are possible. The pathways toward dissipation of the dominant geostrophic eddy KE depend crucially on the direction of the cascade but are difficult to quantify because of serious observational difficulties for wavelengths shorte...

Ocean Circulation Kinetic Energy: Reservoirs, Sources, and Sinks

. The Canadian Earth System Model version 5 (CanESM5) is a global
model developed to simulate historical climate change and variability, to
make centennial-scale projections of future climate, and to produce
initialized seasonal and decadal predictions. This paper describes the model
components and their coupling, as well as various aspects of model
development, including tuning, optimization, and a reproducibility strategy.
We also document the stability of the model using a long control simulation,
quantify the model's ability to reproduce large-scale features of the
historical climate, and evaluate the response of the model to external
forcing. CanESM5 is comprised of three-dimensional atmosphere (T63 spectral
resolution equivalent roughly to 2.8 ∘ ) and ocean (nominally 1 ∘ ) general
circulation models, a sea-ice model, a land surface scheme, and explicit
land and ocean carbon cycle models. The model features relatively coarse
resolution and high throughput, which facilitates the production of large
ensembles. CanESM5 has a notably higher equilibrium climate sensitivity
(5.6 K) than its predecessor, CanESM2 (3.7 K), which we briefly discuss, along
with simulated changes over the historical period. CanESM5 simulations
contribute to the Coupled Model Intercomparison Project phase 6 (CMIP6)
and will be employed for climate science and service applications in Canada.

/pdf/the-canadian-earth-system-model-version-5-canesm5-0-3-3yhcmcmmeu.pdf

The Canadian Earth System Model version 5 (CanESM5.0.3)

If model parameterizations of unresolved physics, such as the variety of upper ocean mixing processes, are to hold over the large range of time and space scales of importance to climate, they must be strongly physically based. Observations, theories, and models of oceanic vertical mixing are surveyed. Two distinct regimes are identified: ocean mixing in the boundary layer near the surface under a variety of surface forcing conditions (stabilizing, destabilizing, and wind driven), and mixing in the ocean interior due to internal waves, shear instability, and double diffusion (arising from the different molecular diffusion rates of heat and salt). Mixing schemes commonly applied to the upper ocean are shown not to contain some potentially important boundary layer physics. Therefore a new parameterization of oceanic boundary layer mixing is developed to accommodate some of this physics. It includes a scheme for determining the boundary layer depth h, where the turbulent contribution to the vertical shear of a bulk Richardson number is parameterized. Expressions for diffusivity and nonlocal transport throughout the boundary layer are given. The diffusivity is formulated to agree with similarity theory of turbulence in the surface layer and is subject to the conditions that both it and its vertical gradient match the interior values at h. This nonlocal “K profile parameterization” (KPP) is then verified and compared to alternatives, including its atmospheric counterparts. Its most important feature is shown to be the capability of the boundary layer to penetrate well into a stable thermocline in both convective and wind-driven situations. The diffusivities of the aforementioned three interior mixing processes are modeled as constants, functions of a gradient Richardson number (a measure of the relative importance of stratification to destabilizing shear), and functions of the double-diffusion density ratio, Rρ. Oceanic simulations of convective penetration, wind deepening, and diurnal cycling are used to determine appropriate values for various model parameters as weak functions of vertical resolution. Annual cycle simulations at ocean weather station Papa for 1961 and 1969–1974 are used to test the complete suite of parameterizations. Model and observed temperatures at all depths are shown to agree very well into September, after which systematic advective cooling in the ocean produces expected differences. It is argued that this cooling and a steady salt advection into the model are needed to balance the net annual surface heating and freshwater input. With these advections, good multiyear simulations of temperature and salinity can be achieved. These results and KPP simulations of the diurnal cycle at the Long-Term Upper Ocean Study (LOTUS) site are compared with the results of other models. It is demonstrated that the KPP model exchanges properties between the mixed layer and thermocline in a manner consistent with observations, and at least as well or better than alternatives.

Oceanic vertical mixing: a review and a model with a nonlocal boundary layer parameterization

This is a review about the Atlantic Meridional Overturning Circulation (AMOC), its mean structure, temporal variability, controlling mechanisms, and role in the coupled climate system. The AMOC plays a central role in climate through its heat and freshwater transports. Northward ocean heat transport achieved by the AMOC is responsible for the relative warmth of the Northern Hemisphere compared to the Southern Hemisphere and is thought to play a role in setting the mean position of the Intertropical Convergence Zone north of the equator. The AMOC is a key means by which heat anomalies are sequestered into the ocean's interior and thus modulates the trajectory of climate change. Fluctuations in the AMOC have been linked to low-frequency variability of Atlantic sea surface temperatures with a host of implications for climate variability over surrounding landmasses. On intra-annual timescales, variability in AMOC is large and primarily reflects the response to local wind forcing; meridional coherence of anomalies is limited to that of the wind field. On interannual to decadal timescales, AMOC changes are primarily geostrophic and related to buoyancy anomalies on the western boundary. A pacemaker region for decadal AMOC changes is located in a western “transition zone” along the boundary between the subtropical and subpolar gyres. Decadal AMOC anomalies are communicated meridionally from this region. AMOC observations, as well as the expanded ocean observational network provided by the Argo array and satellite altimetry, are inspiring efforts to develop decadal predictability systems using coupled atmosphere-ocean models initialized by ocean data.

https://agupubs.onlinelibrary.wiley.com/doi/pdf/10.1002/2015RG000493

Observations, inferences, and mechanisms of the Atlantic Meridional Overturning Circulation: A review

Ocean eddies generated through instability of the mean flow are a vital component of the energy budget of the global ocean. Modelling combined with satellite altimetry data suggests that the energy from westward-propagating eddies is scattered and eventually dispersed when they reach the western boundary of an ocean basin. Ocean eddies generated through instability of the mean flow are a vital component of the energy budget of the global ocean1,2,3. In equilibrium, the sources and sinks of eddy energy have to be balanced. However, where and how eddy energy is removed remains uncertain3,4. Ocean eddies are observed to propagate westwards at speeds similar to the phase speeds of classical Rossby waves5, but what happens to the eddies when they encounter the western boundary is unclear. Here we use a simple reduced-gravity model along with satellite altimetry data to show that the western boundary acts as a ‘graveyard’ for the westward-propagating ocean eddies. We estimate a convergence of eddy energy near the western boundary of approximately 0.1–0.3 TW, poleward of 10° in latitude. This energy is most probably scattered into high-wavenumber vertical modes, resulting in energy dissipation and diapycnal mixing6. If confirmed, this eddy-energy sink will have important implications for the ocean circulation.

/pdf/significant-sink-of-ocean-eddy-energy-near-western-14uz99rx9x.pdf

Significant sink of ocean-eddy energy near western boundaries

Wind-induced near-inertial energy has been believed to be an important source for generating the ocean mixing required to maintain the global meridional overturning circulation. In the present study, the near-inertial energy budget in a realistic (1)/(12)degrees model of the North Atlantic Ocean driven by synoptically varying wind forcing is examined. The authors find that nearly 70% of the wind-induced near-inertial energy at the sea surface is lost to turbulent mixing within the top 200 m and, hence, is not available to generate diapycnal mixing at greater depth. Assuming this result can be extended to the global ocean, it is estimated that the wind-induced near-inertial energy available for ocean mixing at depth is, at most, 0.1 TW. This confirms a recent suggestion that the role of wind-induced near-inertial energy in sustaining the global overturning circulation might have been overemphasized.

/pdf/on-the-loss-of-wind-induced-near-inertial-energy-to-3bqex7nsvf.pdf

On the Loss of Wind-Induced Near-Inertial Energy to Turbulent Mixing in the Upper Ocean

The interaction between inertial oscillations generated by a storm and a mesoscale eddy field is studied using a Southern Ocean channel model. It is shown that the leakage of near-inertial energy out of the surface layer is strongly enhanced by the presence of the eddies, with the anticyclonic eddies acting as a conduit to the deep ocean. Given the ubiquity of the atmospheric storm tracks (a source of near-inertial energy for the ocean) and regions of strong ocean mesoscale variability, we argue that this effect could be important for understanding pathways by which near-inertial energy enters the ocean and is ultimately available for mixing.

/pdf/enhanced-vertical-propagation-of-storm-induced-near-inertial-i1dvccfm4t.pdf

Enhanced vertical propagation of storm-induced near-inertial energy in an eddying ocean channel model

The work done by the wind over the northwest Atlantic Ocean is examined using a realistic high-resolution ocean model driven by synoptic wind forcing. Two model runs are conducted with the difference only in the way the wind stress is calculated. Our results show that the effect of including ocean surface currents in the wind stress formulation is to reduce the total wind work integrated over the model domain by about 17%. The reduction is caused by a sink term in the wind work calculation associated with the presence of ocean currents. In addition, the modelled eddy kinetic energy decreases by about 10%, in response to direct mechanical damping by the surface stress. A simple scaling argument shows that the latter can be expected to be more important than bottom friction in the energy budget.

/pdf/wind-work-in-a-model-of-the-northwest-atlantic-ocean-4rj7o17hgi.pdf

Wind work in a model of the northwest Atlantic Ocean

The wind power input to the ocean general circulation is usually calculated from the time-averaged wind products. Here, this wind power input is reexamined using available observations, focusing on the role of the synoptically varying wind. Power input to the ocean general circulation is found to increase by over 70% when 6-hourly winds are used instead of monthly winds. Much of the increase occurs in the storm-track regions of the Southern Ocean, Gulf Stream, and Kuroshio Extension. This result holds irrespective of whether the ocean surface velocity is accounted for in the wind stress calculation. Depending on the fate of the highfrequency wind power input, the power input to the ocean general circulation relevant to deep-ocean mixing may be less than previously thought. This study emphasizes the difficulty of choosing appropriate forcing for ocean-only models.

http://ocean.mit.edu/~cwunsch/papersonline/zhai_etal_2012_windwork_jpo.pdf

Xiaoming Zhai

Papers

Significant sink of ocean-eddy energy near western boundaries

On the Loss of Wind-Induced Near-Inertial Energy to Turbulent Mixing in the Upper Ocean

Enhanced vertical propagation of storm-induced near-inertial energy in an eddying ocean channel model

Wind work in a model of the northwest Atlantic Ocean

On the Wind Power Input to the Ocean General Circulation