We synthesize data on all known extant and fossil Coleoptera family-group names for the first time. A catalogue of 4887 family-group names (124 fossil, 4763 extant) based on 4707 distinct genera in Coleoptera is given. A total of 4492 names are available, 183 of which are permanently invalid because they are based on a preoccupied or a suppressed type genus. Names are listed in a classification framework. We recognize as valid 24 superfamilies, 211 families, 541 subfamilies, 1663 tribes and 740 subtribes. For each name, the original spelling, author, year of publication, page number, correct stem and type genus are included. The original spelling and availability of each name were checked from primary literature. A list of necessary changes due to Priority and Homonymy problems, and actions taken, is given. Current usage of names was conserved, whenever possible, to promote stability of the classification.

New synonymies (family-group names followed by genus-group names): Agronomina Gistel, 1848 syn. nov. of Amarina Zimmermann, 1832 (Carabidae), Hylepnigalioini Gistel, 1856 syn. nov. of Melandryini Leach, 1815 (Melandryidae), Polycystophoridae Gistel, 1856 syn. nov. of Malachiinae Fleming, 1821 (Melyridae), Sclerasteinae Gistel, 1856 syn. nov. of Ptilininae Shuckard, 1839 (Ptinidae), Phloeonomini Adam, 2001 syn. nov. of Omaliini MacLeay, 1825 (Staphylinidae), Sepedophilini Adam, 2001 syn. nov. of Tachyporini MacLeay, 1825 (Staphylinidae), Phibalini Gistel, 1856 syn. nov. of Cteniopodini Solier, 1835 (Tenebrionidae); Agronoma Gistel 1848 (type species Carabus familiaris Duftschmid, 1812, designated herein) syn. nov. of Amara Bonelli, 1810 (Carabidae), Hylepnigalio Gistel, 1856 (type species Chrysomela caraboides Linnaeus, 1760, by monotypy) syn. nov. of Melandrya Fabricius, 1801 (Melandryidae), Polycystophorus Gistel, 1856 (type species Cantharis aeneus Linnaeus, 1758, designated herein) syn. nov. of Malachius Fabricius, 1775 (Melyridae), Sclerastes Gistel, 1856 (type species Ptilinus costatus Gyllenhal, 1827, designated herein) syn. nov. of Ptilinus Geoffroy, 1762 (Ptinidae), Paniscus Gistel, 1848 (type species Scarabaeus fasciatus Linnaeus, 1758, designated herein) syn. nov. of Trichius Fabricius, 1775 (Scarabaeidae), Phibalus Gistel, 1856 (type species Chrysomela pubescens Linnaeus, 1758, by monotypy) syn. nov. of Omophlus Dejean, 1834 (Tenebrionidae). The following new replacement name is proposed: Gompeliina Bouchard, 2011 nom. nov. for Olotelina Baguena Corella, 1948 (Aderidae).

Reversal of Precedence (Article 23.9) is used to conserve usage of the following names (family-group names followed by genus-group names): Perigonini Horn, 1881 nom. protectum over Trechicini Bates, 1873 nom. oblitum (Carabidae), Anisodactylina Lacordaire, 1854 nom. protectum over Eurytrichina LeConte, 1848 nom. oblitum (Carabidae), Smicronychini Seidlitz, 1891 nom. protectum over Desmorini LeConte, 1876 nom. oblitum (Curculionidae), Bagoinae Thomson, 1859 nom. protectum over Lyprinae Gistel 1848 nom. oblitum (Curculionidae), Aterpina Lacordaire, 1863 nom. protectum over Heliomenina Gistel, 1848 nom. oblitum (Curculionidae), Naupactini Gistel, 1848 nom. protectum over Iphiini Schonherr, 1823 nom. oblitum (Curculionidae), Cleonini Schonherr, 1826 nom. protectum over Geomorini Schonherr, 1823 nom. oblitum (Curculionidae), Magdalidini Pascoe, 1870 nom. protectum over Scardamyctini Gistel, 1848 nom. oblitum (Curculionidae), Agrypninae/-ini Candeze, 1857 nom. protecta over Adelocerinae/-ini Gistel, 1848 nom. oblita and Pangaurinae/-ini Gistel, 1856 nom. oblita (Elateridae), Prosternini Gistel, 1856 nom. protectum over Diacanthini Gistel, 1848 nom. oblitum (Elateridae), Calopodinae Costa, 1852 nom. protectum over Sparedrinae Gistel, 1848 nom. oblitum (Oedemeridae), Adesmiini Lacordaire, 1859 nom. protectum over Macropodini Agassiz, 1846 nom. oblitum (Tenebrionidae), Bolitophagini Kirby, 1837 nom. protectum over Eledonini Billberg, 1820 nom. oblitum (Tenebrionidae), Throscidae Laporte, 1840 nom. protectum over Stereolidae Rafinesque, 1815 nom. oblitum (Throscidae) and Lophocaterini Crowson, 1964 over Lycoptini Casey, 1890 nom. oblitum (Trogossitidae); Monotoma Herbst, 1799 nom. protectum over Monotoma Panzer, 1792 nom. oblitum (Monotomidae); Pediacus Shuckard, 1839 nom. protectum over Biophloeus Dejean, 1835 nom. oblitum (Cucujidae), Pachypus Dejean, 1821 nom. protectum over Pachypus Billberg, 1820 nom. oblitum (Scarabaeidae), Sparrmannia Laporte, 1840 nom. protectum over Leocaeta Dejean, 1833 nom. oblitum and Cephalotrichia Hope, 1837 nom. oblitum (Scarabaeidae).

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Family-group names in Coleoptera (Insecta)

A catalogue of the subfamily Lamiinae (Coleoptera: Cerambycidae) of the Neotropical region is presented. The tribes (36), genera (723), species (4231) and subspecies (60) are listed in alphabetical order. Under each family-group name bibliographical references are given and under each species-group name, data on the type-locality, the acronym of the institution where the type is deposited, the geographical distribution and detailed bibliographical references are provided. Two new combinations are proposed: Urographis eucharis (Bates, 1885), new comb. and Urographis vexillaris (Bates, 1872) new comb., both from Graphisurus LeConte, 1852, not Kirby, 1837. The following new names are given: Alcathousiella new name to replace Alcathous Thomson, 1864, preoccupied by Alcathous Stal, 1863, Hemiptera; Camposiellina new name to replace Camposiella Lane, 1972, preoccupied by Camposiella Hebard, 1924, Orthoptera; Edechthistatus new name to replace Parechthistatus Giesbert, 2001, preoccupied by Parechthistatus Breuning, 1942, Coleoptera; Elytracanthina new name to replace Elytracantha Lane, 1955, preoccupied by Elytracantha Kleine, 1915, Coleoptera; Eranina new name to replace Erana Bates, 1866, preoccupied by Erana Gray, 1840, Aves; Heteresmia new name to replace Esmia Pascoe, 1859, preoccupied by Esmia Leach, 1847, Mollusca; Eupalessa new name to replace Eupales Dillon & Dillon, 1945, preoccupied by Eupales Lefevre, 1885, Coleoptera; Melzerus new name to replace Idiomerus Melzer, 1934, preoccupied by Idiomerus Imms, 1912, Collembola; Midamiella new name to replace Midamus Dillon & Dillon, 1945, preoccupied by Midamus Simon, 1881, Arachnida; Neoamphion new name to replace Amphion Reiche, 1840, preoccupied by Amphion Huebner, 1819, Lepidoptera; Neocolobura new name to replace Colobura Blanchard, 1851, preoccupied by Colobura Billberg, 1820, Lepidoptera; Neohoplonotus new name to replace Hoplonotus Blanchard, 1851, preoccupied by Hoplonotus Schmidt-Goebel, 1846, Coleoptera; Neohylus new name to replace Hylus Dillon & Dillon, 1945, preocuppied by Hylus Van Dyke, 1945, Coleoptera; Neolampedusa new name to replace Lampedusa Dillon & Dillon, 1945, preoccupied by Lampedusa Boettger, 1877, Mollusca; Proseriphus new name to replace Seriphus Bates, 1864, preoccupied by Seriphus Ayres, 1857, Pisces. One new synonym is proposed: Proxepectasis Monne & Giesbert, 1992 = Parepectasoides Breuning, 1979.

Catalogue of the Cerambycidae (Coleoptera) of the Neotropical Region. Part II. Subfamily Lamiinae

A large-scale phylogenetic study is presented for Cucujoidea (Coleoptera), a diverse superfamily of beetles that historically has been taxonomically difficult. This study is the most comprehensive analysis of cucujoid taxa to date, with DNA sequence data sampled from eight genes (four nuclear, four mitochondrial) for 384 coleopteran taxa, including exemplars of 35 (of 37) families and 289 genera of Cucujoidea. Maximum-likelihood analyses of these data present many significant relationships, some proposed previously and some novel. Tenebrionoidea and Lymexyloidea are recovered together and Cleroidea forms the sister group to this clade. Chrysomeloidea and Curculionoidea are recovered as sister taxa and this clade (Phytophaga) forms the sister group to the core Cucujoidea (Cucujoidea s.n.). The nitidulid series is recovered as the earliest-diverging core cucujoid lineage, although the earliest divergences among core Cucujoidea are only weakly supported. The cerylonid series (CS) is recovered as monophyletic and is supported as a major Cucujiform clade, sister group to the remaining superfamilies of Cucujiformia. Currently recognized taxa that were not recovered as monophyletic include Cucujoidea, Endomychidae, Cerylonidae and Bothrideridae. Biphyllidae and Byturidae were recovered in Cleroidea. The remaining Cucujoidea were recovered in two disparate major clades: one comprising the nitidulid series + erotylid series + Boganiidae and Hobartiidae + cucujid series, and the other comprising the cerylonid series. Propalticidae are recovered within Laemophloeidae. The cerylonid series includes two major clades, the bothriderid group and the coccinellid group. Akalyptoischiidae are recovered as a separate clade from Latridiidae. Eupsilobiinae are recovered as the sister taxon to Coccinellidae. In light of these findings, many formal changes to cucujiform beetle classification are proposed. Biphyllidae and Byturidae are transferred to Cleroidea. The cerylonid series is formally recognized as a new superfamily, Coccinelloidea stat.n. Current subfamilies elevated (or re-elevated) to family status include: Murmidiidae stat.n., Teredidae stat.n., Euxestidae stat.n., Anamorphidae stat.rev., Eupsilobiidae stat.n., and Mycetaeidae stat.n. The following taxa are redefined and characterized: Cleroidea s.n., Cucujoidea s.n., Cerylonidae s.n., Bothrideridae s.n., Endomychidae s.n. A new subfamily, Cyclotominae stat.n., is described. Stenotarsinae syn.n. is formally subsumed within a new concept of Endomychinae s.n.

Phylogeny and classification of Cucujoidea and the recognition of a new superfamily Coccinelloidea (Coleoptera: Cucujiformia)

A checklist of longhorn beetles (Coleoptera: Cerambycidae) within the present geographical frontier of Indian subcontinent up to 2016 is provided. As per the current checklist prepared, there are 1536 species, classified under 440 genera, 72 tribes, and seven subfamilies of Cerambycidae (Parandrinae is not present in India). The report is accounted for 4.2 per cent of species, 7.94 per cent of genera and 28.24 per cent of tribes from India as compared to global record. For each species, accepted nomenclature followed by all relevant works reporting systematics, distribution and ecology of Indian longhorn beetles is provided along with synonyms, type locality and distribution within and outside India.

The checklist of longhorn beetles (Coleoptera: Cerambycidae) from India

An analysis of the species diversity and distribution of agriculturally important longhorn beetles (cerambycidae: coleoptera) from india

Family-group names proposed for beetles belonging to the family Cerambycidae are catalogued and their availability is determined using the rules of the current International Code of Zoological Nomenclature. A synoptic classification of the family summarizes the validity of the names. Type genera of all family-group names are listed and the type species and stems of genera of available family-group names are included. A new family-group name, Elytracanthinini Bousquet (type genus: Elytracanthina Monn, 2005, a replacement name for Elytracantha Lane, 1955) is proposed for Elytracanthinae Lane, 1955. Ichthyosoma armatum Montrouzier, 1855 is designated as type species of Icthyosoma Boisduval, 1835. Reversal of precedence is used to preserve the validity of the following family-group names: Anaglyptides Lacordaire, 1868 (over Anaglyptisidae Gistel, 1848 [Buprestidae]); Dryobiini Arnett, 1962 (over Dryobiadae Gistel, 1856 [Ptinidae]); Hemilophitae Thomson, 1868 (over Amphionychitae Thomson, 1860) and Heteropsides Lacordaire, 1869 (over Dichophyiaeidae Gistel, 1848). The following family-group names, although junior synonyms, are preserved as valid until an application is submitted to the International Commission on Zoological Nomenclature; in these cases a reversal of precedence could not be applied: Eurypodini Gahan, 1906 (over Zaracinae Pascoe, 1869); Macronides Lacordaire, 1868 (over Enchapteritae Thomson, 1861); Pyresthides Lacordaire, 1868 (over Pseudolepturitae Thomson, 1861 and Erythrinae Pascoe, 1866) and Stenoderinae Pascoe, 1867 (over Syllitae Thomson, 1864). A total of 238 valid cerambycid family-group names (413 available names) are recognized in the following 13 subfamilies: Vesperinae (1 valid family-group name), Oxypeltinae (1), Disteniinae (4), Anoplodermatinae (3), Philinae (1), Parandrinae (2), Prioninae (24), Spondylidinae (5), Necydalinae (1), Lepturinae (8), Lamiinae (80), Dorcasominae (1), and Cerambycinae (107).

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Catalogue of family-group names in Cerambycidae (Coleoptera)

A list of the Cerambycidae of Bolivia is presented. It totals 1,259 species including 496 new country records. When available the known geographical distribution by department is shown for each species.

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Preliminary checklist of Bolivian Cerambycidae (Coleoptera)

A revision of the American species of Prionus Geoffroy, 1762 is presented. Prionus (Neopolyarthron) Semenov, 1899 and Prionus (Antennalia) Casey, 1912 are synonymized with Prionus Geoffroy, 1762. Homaesthesis LeConte, 1873 is considered a true subgenus of Prionus. Prionus (Homaesthesis) rhodocerus Linsley, 1957 and Prionus (Homaesthesis) linsleyi Hovore, 1981 are synonymized with Prionus simplex (Casey, 1912). Prionus beauvoisi Lameere, 1915 and Prionus (Neopolyarthron) debilis Casey, 1924 are synonymized with P. imbricornis (Linnaeus, 1767). Prionus (Neopolyarthron) townsendi Casey, 1912 and Prionus (Neopolyarthron) curticollis Casey, 1912 are synonymized with Prionus mexicanus Bates, 1884. Prionus batesi Lameere, 1920 is synonymized with Prionus aztecus Casey, 1912. Prionus hintoni Linsley, 1935 is synonymized with Prionus flohri Bates, 1884. Prionus (Antennalia) fissicornis parviceps Casey, 1912 is excluded as the synonym of Prionus fissicornis Haldeman, 1846 and instead synonymized with P. imbricornis (Linnaeus, 1767). Prionus (Prionus) validiceps Casey, 1912 is excluded from the synonymy of P. pocularis Dalman, 1817, and synonymized with P. (P.) californicus Motschulsky, 1845. Prionus (Prionus) tumidus Casey, 1912 is excluded from the synonymy of P. heroicus Semenov, 1907, and synonymized with P. (P.) californicus. The lectotype female and the paralectotype male of Prionus (Prionus) tristis are excluded from the synonym of P. (P.) heroicus and transferred to the synonym of P. (P.) californicus; the paralectotype female of P. (P.) tristis is maintained in the synonymy of P. (P.) heroicus. Prionus (Prionus) fontinalis Casey, 1914 is excluded from the synonymy of P. (P.) heroicus and synonymized with P. (P.) californicus. Prionus simplex is formally excluded from the Cerambycidae fauna of Oklahoma, USA. Comments on the page, plate, and figure of publication of Cerambyx laticollis Drury, 1773 are presented. Prionus (Trichoprionus) Fragoso & Monne, 1982 is considered a genus different from Prionus. Hypoprionus is designated as a replacement name for Prionellus Casey, 1924 and Cerambyx laticollis Drury, 1773 is designated as the type species. Comments on the type localities of Prionus emarginatus, Prionus palparis Say, 1824, and Prionus (Neopolyarthron) aztecus Casey, 1912 are presented. Prionus (Homaesthesis) integer sensu Linsley (1962) and Chemsak (1996) is described as P. (H.) geminus, new species. Comments on the date of publication of Prionus fissicornis Haldeman, 1846 are presented. Comments on the status of the syntypes of Cerambyx imbricornis Linnaeus, 1767 are also presented. Lectotype specimens for Prionus flohri Bates, 1884, Prionus (Prionus) tristis, and Prionus lecontei Lameere, 1912 are designated. Comments on the number of specimens used in the original description of Prionus californicus are presented, and a lectotype for this species is designated. New state records are presented for Prionus emarginatus Say, 1824; P. imbricornis (Linnaeus, 1767); P. aztecus Casey, 1912; P. poultoni Lameere, 1912; P. pocularis Dalman, 1817; and P. mexicanus Bates, 1884.

Revision of the American species of the genus Prionus Geoffroy, 1762 (Coleoptera, Cerambycidae, Prioninae, Prionini).

An updated checklist of the Cerambycidae of Costa Rica is presented. This new version includes 1,071 species and subspecies in 429 genera, 69 tribes, and six subfamilies. Of these, 181 are new country records and 136 species are known only from Costa Rica. In addition, provincial distribution data are provided for each species. The checklist supports a wealth of scientific literature in many other groups of flora and fauna indicating Costa Rica has high species richness of cerambycid beetles.

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Checklist of the Cerambycidae (Coleoptera) of Costa Rica

Australophanus, new genus, is described and illustrated for Cryptamorpha redtenbacheri (Reitter). Platamus Erichson is synonymized under Telephanus Erichson, new synonymy. Euplatamus Sharp, new status, replaces Platamus Erichson as the genus name. Type species are designated for Aplatamus Grouvelle and Euplatamus Sharp. Telephanus velox (Haldeman) is synonymized under Telephanus atricapillus Erichson. A diagnosis of the tribe Telephanini, a key to the described telephanine genera of the world is presented, and a phylogeny of the family Silvanidae is proposed.

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Eugenio H. Nearns

Papers

Catalogue of family-group names in Cerambycidae (Coleoptera)

Preliminary checklist of Bolivian Cerambycidae (Coleoptera)

Revision of the American species of the genus Prionus Geoffroy, 1762 (Coleoptera, Cerambycidae, Prioninae, Prionini).

Checklist of the Cerambycidae (Coleoptera) of Costa Rica

0048. A new genus of telephanine Silvanidae (Coleoptera: Cucujoidea), with a diagnosis of the tribe and key to genera