抗原变异可使得多种致病微生物易于逃避宿主免疫应答。表达在感染红细胞表面的恶性疟原虫红细胞表面蛋白1（PfPMP1）与感染红细胞、内皮细胞、树突状细胞以及胎盘的单个或多个受体作用，在黏附及免疫逃避中起关键的作用。每个单倍体基因组var基因家族编码约60种成员，通过启动转录不同的var基因变异体为抗原变异提供了分子基础。

疟原虫var基因转换速率变化导致抗原变异[英]／Paul H, Robert P, Christodoulou Z, et al//Proc Natl Acad Sci U S A

The terrestrial carbon sink has been large in recent decades, but its size and location remain uncertain. Using forest inventory data and long-term ecosystem carbon studies, we estimate a total forest sink of 2.4 ± 0.4 petagrams of carbon per year (Pg C year–1) globally for 1990 to 2007. We also estimate a source of 1.3 ± 0.7 Pg C year–1 from tropical land-use change, consisting of a gross tropical deforestation emission of 2.9 ± 0.5 Pg C year–1 partially compensated by a carbon sink in tropical forest regrowth of 1.6 ± 0.5 Pg C year–1. Together, the fluxes comprise a net global forest sink of 1.1 ± 0.8 Pg C year–1, with tropical estimates having the largest uncertainties. Our total forest sink estimate is equivalent in magnitude to the terrestrial sink deduced from fossil fuel emissions and land-use change sources minus ocean and atmospheric sinks.

/pdf/a-large-and-persistent-carbon-sink-in-the-world-s-forests-2rgrfjaqnh.pdf

A Large and Persistent Carbon Sink in the World’s Forests

https://www.nrs.fs.fed.us/pubs/jrnl/2011/nrs_2011_pan_002-supporting.pdf

Supporting Online Material for A Large and Persistent Carbon Sink in the World's Forests

United States Agency for International Development

Forest systems cover more than 4.1 x 109 hectares of the Earth9s land area. Globally, forest vegetation and soils contain about 1146 petagrams of carbon, with approximately 37 percent of this carbon in low-latitude forests, 14 percent in mid-latitudes, and 49 percent at high latitudes. Over two-thirds of the carbon in forest ecosystems is contained in soils and associated peat deposits. In 1990, deforestation in the low latitudes emitted 1.6 ± 0.4 petagrams of carbon per year, whereas forest area expansion and growth in mid- and high-latitude forest sequestered 0.7 ± 0.2 petagrams of carbon per year, for a net flux to the atmosphere of 0.9 ± 0.4 petagrams of carbon per year. Slowing deforestation, combined with an increase in forestation and other management measures to improve forest ecosystem productivity, could conserve or sequester significant quantities of carbon. Future forest carbon cycling trends attributable to losses and regrowth associated with global climate and land-use change are uncertain. Model projections and some results suggest that forests could be carbon sinks or sources in the future.

https://science.sciencemag.org/content/sci/263/5144/185.full.pdf

Carbon pools and flux of global forest ecosystems.

Estimates of national-scale forest carbon (C) stocks and fluxes are typically based on allometric regression equations developed using dimensional analysis techniques. However, the literature is inconsistent and incomplete with respect to large-scale forest C estimation. We compiled all available diameter-based allometric regression equations for estimating total aboveground and component biomass, defined in dry weight terms, for trees in the United States. We then implemented a modified meta-analysis based on the published equations to develop a set of consistent, national-scale aboveground biomass regression equations for U.S. species. Equations for predicting biomass of tree components were developed as proportions of total aboveground biomass for hardwood and softwood groups. A comparison with recent equations used to develop large-scale biomass estimates from U.S. forest inventory data for eastern U.S. species suggests general agreement (±30%) between biomass estimates. The comparison also shows that differences in equation forms and species groupings may cause differences at small scales depending on tree size and forest species composition. This analysis represents the first major effort to compile and analyze all available biomass literature in a consistent national-scale framework. The equations developed here are used to compute the biomass estimates used by the model FORCARB to develop the U.S. C budget. FOR. SCI. 49(1):12-35.

https://www.fs.fed.us/ne/newtown_square/publications/other_publishers/OCR/ne_2003jenkins01.pdf

National scale biomass estimators for United States tree species

Forests are the dominant terrestrial ecosystem on Earth. We review the environmental factors controlling their structure and global distribution and evaluate their current and future trajectory. Adaptations of trees to climate and resource gradients, coupled with disturbances and forest dynamics, create complex geographical patterns in forest assemblages and structures. These patterns are increasingly discernible through new satellite and airborne observation systems, improved forest inventories, and global ecosystem models. Forest biomass is a complex property affected by forest distribution, structure, and ecological processes. Since at least 1990, biomass density has consistently increased in global established forests, despite increasing mortality in some regions, suggesting that a global driver such as elevated CO2 may be enhancing biomass gains. Global forests have also apparently become more dynamic. Advanced information about the structure, distribution, and biomass of the world's forests provides...

/pdf/the-structure-distribution-and-biomass-of-the-world-s-674p60qv9w.pdf

The Structure, Distribution, and Biomass of the World's Forests

Carbon accumulation in forests has been attributed to historical changes in land use and the enhancement of tree growth by CO 2 fertilization, N deposition, and climate change. The relative contribution of land use and growth enhancement is estimated by using inventory data from five states spanning a latitudinal gradient in the eastern United States. Land use is the dominant factor governing the rate of carbon accumulation in these states, with growth enhancement contributing far less than previously reported. The estimated fraction of aboveground net ecosystem production due to growth enhancement is 2.0 ± 4.4%, with the remainder due to land use.

https://science.sciencemag.org/content/sci/290/5494/1148.full.pdf

Contributions of Land-Use History to Carbon Accumulation in U.S. Forests

A database consisting of 2,640 equations compiled from the literature for predicting the biomass of trees and tree components from diameter measurements of species found in North America. Bibliographic information, geographic locations, diameter limits, diameter and biomass units, equation forms, statistical errors, and coefficients are provided for each equation, along with examples of how to use the database. The CD-ROM included with the paper version of this publication contains the complete database (Table 3) in spreadsheet format (Microsoft Excel 2002� with Windows XP�). The database files can also be viewed in both spreadsheet and pdf formats by directing your browser to the Global Change page at http://www.fs.fed.us/ne/global/pubs/books/index.html

https://www.fs.fed.us/ne/newtown_square/publications/technical_reports/pdfs/2004/ne_gtr319.pdf

Richard A. Birdsey

Papers

Supporting Online Material for A Large and Persistent Carbon Sink in the World's Forests

National scale biomass estimators for United States tree species

The Structure, Distribution, and Biomass of the World's Forests

Contributions of Land-Use History to Carbon Accumulation in U.S. Forests

Comprehensive database of diameter-based biomass regressions for North American tree species