Showing papers on "Gondwana published in 2022"

Grasses through space and time: An overview of the biogeographical and macroevolutionary history of Poaceae

Abstract: Grasses are widespread on every continent and are found in all terrestrial biomes. The dominance and spread of grasses and grassland ecosystems have led to significant changes in Earth′s climate, geochemistry, and biodiversity. The abundance of DNA sequence data, particularly chloroplast sequences, and advances in placing grass fossils within the family allows for a reappraisal of the family′s origins, timing, and geographic spread and the factors that have promoted diversification. We reconstructed a time‐calibrated grass phylogeny and inferred ancestral areas using chloroplast DNA sequences from nearly 90% of extant grass genera. With a few notable exceptions, the phylogeny is well resolved to the subtribal level. The family began to diversify in the Early–Late Cretaceous (crown age of 98.54 Ma) on West Gondwana before the complete split between Africa and South America. Vicariance from the splitting of Gondwana may be responsible for the initial divergence in the family. However, Africa clearly served as the center of origin for much of the early diversification of the family. With this phylogenetic, temporal, and spatial framework, we review the evolution and biogeography of the family with the aim to facilitate the testing of biogeographical hypotheses about its origins, evolutionary tempo, and diversification. The current classification of the family is discussed with an extensive review of the extant diversity and distribution of species, molecular and morphological evidence supporting the current classification scheme, and the evidence informing our understanding of the biogeographical history of the family.

Reconstructing the Variscan Terranes in the Alpine Basement: Facts and Arguments for an Alpidic Orocline

Abstract: The existence of pieces of the Variscan belt in the Alpine basement has been acknowledged for a long time but the correlation of these massifs to the litho-tectonic domains established in Western Europa outside the Alpine chain is still disputed. Due to their ubiquitous character, the abundant late Variscan migmatites and granites are useless to reconstruct the Variscan architecture in the Alpine basement. Ophiolitic sutures, high- and low-grade metamorphic units, and foreland basins provide a preliminary reconstruction of the Variscan orogen exposed in the Alpine basement. The longitudinal extension of the Armorican and Saxo-Thuringian microcontinents between Laurussia and Gondwana is proposed independently of the Intra-alpine and Galatian terranes. The litho-tectonic units of the Corsica-Sardinia segment are correlated to the Moldanubian, Armorican and Saxo-Thuringian Domains. In the Alpine Helvetic and Penninic Domains, the Chamrousse ophiolites are ascribed to the Tepla-Le Conquet suture, whereas the Lepontine, and Stubach ophiolites represent the Rheic suture. The south-directed nappe stack of the South Alpine Domain is similar to the Moldanubian French Massif Central. In the Austroalpine nappe stack, the Ritting ophiolites separate Saxo-Thuringia and Armorica continental blocks. Disentangling the Variscan belt in the Alpine basement suggests a concave-to-the-East arcuate structure called here the Variscan Alpidic orocline.

A titanosaurian sauropod with Gondwanan affinities in the latest Cretaceous of Europe

Abstract: The origin of the last sauropod dinosaur communities in Europe and their evolution during the final 15 million years of the Cretaceous have become a complex phylogenetic and palaeobiogeographic puzzle characterized by the controversy on the alleged coexistence of immigrant, Gondwana-related taxa alongside relictual and insular clades. In this context, we describe a new titanosaurian sauropod dinosaur, Abditosaurus kuehnei gen. et sp. nov., from the Late Cretaceous (Maastrichtian) Tremp Group of Catalonia (Spain). Phylogenetic analyses recover Abditosaurus separately from other European titanosaurs, within a clade of otherwise South American and African saltasaurines. The affinity of the new taxon with southern landmasses is reinforced by spatiotemporal co-occurrence with Gondwanan titanosaurian oospecies in southern Europe. The large size and the lack of osteohistological features potentially related to insular dwarfism or size reduction support the idea that Abditosaurus belongs to an immigrant lineage, unequivocally distinct from some of the island dwarfs of the European archipelago. The arrival of the Abditosaurus lineage to the Ibero-Armorican Island is hypothesized to have occurred during the earliest Maastrichtian (70.6 Ma), probably as a result of a global and regional sea-level drop that reactivated ancient dispersal routes between Africa and Europe. The arrival of large-bodied titanosaurs to the European archipelago produced dramatic changes in its insular ecosystems and important evolutionary changes in its dinosaur faunas, especially with respect to the 'island rule' effect.

Strain Partitioning along Terrane Bounding and Intraterrane Shear Zones: Constraints from a Long-Lived Transpressional System in West Gondwana (Ribeira Belt, Brazil)

Abstract: Shear zones are zones of localized high strain accommodating differential motion in the lithosphere and impacting the crustal rheology and deformational history of orogenic belts. Although terrane bounding shear zones are widely studied, intraterrane shear zones and their tectonic significance, especially in association with supercontinent assembly, is a largely unexplored topic. The Ribeira Belt (SE Brazil), a Neoproterozoic-Cambrian orogenic belt from West Gondwana, is dissected by a crustal-scale NE-trending transcurrent shear zone system that juxtaposes composite terranes. Despite its extensive coverage and complexity, this shear zone system remains poorly investigated. In this paper, we explore the thermal and deformational regimes, and timing of ductile shearing using a multiscale approach combining structural analysis derived from remote sensing and field-based structural data, microstructures, quantitative structural analysis, and multimineral U–Pb geochronology (zircon, titanite, monazite, and xenotime). Our data, combined with previously published data, indicate a transitional northeastward increase in metamorphic conditions from lower greenschist to granulite facies conditions (from 250–300 to 750–800°C), reflecting the different crustal levels that are exposed. Vorticity and finite strain data indicate a complex strain regime with varied contributions of pure and simple shear and oblate-shape ellipsoids in strike-slip shear zones and prolate-shaped ellipsoids in dip-slip reverse shear zones. The strain set suggests that all shear zones were developed under subsimple shear deformational regimes involving thrusting and folding followed by wrench tectonics. The pure shear component of deformation was accommodated in folded domains between shear zones. Geochronological data suggest intermittent ductile shear zone activations from ca. 900–830 to 530 Ma, partially coeval with at least two major episodes of terrane accretion at 850–760 Ma and 610–585 Ma. The spatial and temporal record of shear zones within the Ribeira Belt indicates that some relate to assembly of the belt and represent either terrane bounding structures (e.g., Itapirapuã shear zone) or intraterrane structures (e.g., Ribeira, Figueira, and Agudos Grandes shear zones), whereas others are terrane bounding, postcollisional shear zones (e.g., Taxaquara shear zone) reactivated in an intracontinental setting (560–535 Ma).

End-permian burnout: the role of permian–triassic wildfires in extinction, carbon cycling, and environmental change in eastern gondwana

Abstract: Abstract: Wildfire has been implicated as a potential driver of deforestation and continental biodiversity loss during the end-Permian extinction event (EPE; ∼ 252 Ma). However, it cannot be established whether wildfire activity was anomalous during the EPE without valid pre- and post-EPE baselines. Here, we assess the changes in wildfire activity in the high-latitude lowlands of eastern Gondwana by presenting new long-term, quantitative late Permian (Lopingian) to Early Triassic records of dispersed fossil charcoal and inertinite from sediments of the Sydney Basin, eastern Australia. We also document little-transported fossil charcoal occurrences in middle to late Permian (Guadalupian to Lopingian) permineralized peats of the Lambert Graben, East Antarctica, and Sydney and Bowen basins, eastern Australia, indicating that even vegetation of consistently moist high-latitude settings was prone to regular fire events. Our records show that wildfires were consistently prevalent through the Lopingian, but the EPE demonstrates a clear spike in activity. The relatively low charcoal and inertinite baseline for the Early Triassic is likely due in part to the lower vegetation density, which would have limited fire spread. We review the evidence for middle Permian to Lower Triassic charcoal in the geosphere, and the impacts of wildfires on sedimentation processes and the evolution of landscapes. Moreover, we assess the evidence of continental extinction drivers during the EPE within eastern Australia, and critically evaluate the role of wildfires as a cause and consequence of ecosystem collapse. The initial intensification of the fire regime during the EPE likely played a role in the initial loss of wetland carbon sinks, and contributed to increased greenhouse gas emissions and land and freshwater ecosystem changes. However, we conclude that elevated wildfire frequency was a short-lived phenomenon; recurrent wildfire events were unlikely to be the direct cause of the subsequent long-term absence of peat-forming wetland vegetation, and the associated ‘coal gap’ of the Early Triassic.

From Gondwana to the Yellow Sea, evolutionary diversifications of true toads Bufo sp. in the Eastern Palearctic and a revisit of species boundaries for Asian lineages

Abstract: Taxa with vast distribution ranges often display unresolved phylogeographic structures and unclear taxonomic boundaries resulting in hidden diversity. This hypothesis-driven study reveals the evolutionary history of Bufonidae, covering the phylogeographic patterns found in Holarctic bufonids from the West Gondwana to the phylogenetic taxonomy of Asiatic true toads in the Eastern Palearctic. We used an integrative approach relying on fossilized birth-death calibrations, population dynamics, gene-flow, species distribution, and species delimitation modeling to resolve the biogeography of the clade and highlight cryptic lineages. We verified the near-simultaneous Miocene radiations within Western and Eastern Palearctic Bufo, c. 14.49-10.00 Mya, temporally matching with the maximum dust outflows in Central Asian deserts. Contrary to earlier studies, we demonstrated that the combined impacts of long dispersal and ice-age refugia equally contributed to the current genetic structure of Bufo in East Asia. Our findings reveal a climate-driven adaptation in septentrional Eastern Asian Bufo, explaining its range shifts toward northern latitudes. We resolve species boundaries within the Eastern Palearctic Bufo, and redefine the taxonomic and conservation units of the northeastern species: B. sachalinensis and its subspecies.The east Asian Asiatic toad (also known by the latin name Bufo gargarizans) lives in a wide range of habitats across East Asia including forests, meadows and cultivated land. However, it remains unclear how these toads evolved and became so widespread – partly because it has proved difficult for researchers to clearly define the species and what distinguishes it from other closely-related species of toads (collectively known as Bufo toads). Othman et al. combined several bioinformatics techniques to study Asiatic toads and 38 other species of bufonid toads from across the globe. This approach found that Bufo toads first emerged in eastern Asia between 14 and 10 million years ago. This coincides with a point in time when large swathes of land in central Asia turned from adequate to sustain toad populations into desert, suggesting the change in climate prompted the toads to migrate eastwards from central Asia. The Bufo toads then divided into two groups of species: one in mainland East Asia and the other in Japan. Furthermore, the study revealed there is more genetic diversity – that is, more variety in the DNA of individuals – in Asiatic toads than previously thought. The findings also help to define several other species of Bufo toads more clearly and describe a new toad species restricted to the Korean Peninsula, northeastern China and the Russian Primorye region: the Sakhalin toad (Bufo sachalinensis). This work demonstrates that a large-scale study of many species across the globe can be used to understand how the species evolved and more clearly distinguish one species from another. The findings of Othman et al. will be of interest to both professional and citizen scientists interested in the natural history of Asia. Furthermore, as several species of Bufo toads are in decline in the wild, they provide evidence that may aid future efforts to conserve them.

Charophytes from the Cretaceous–Palaeogene transition in the Jhilmili intertrappean beds of Central India

Abstract: The Jhilmili intertrappean beds (Chhindwara District, Madhya Pradesh) of Central India are studied from a palaeoenvironmental, palaeoecological, and palaeobiogeographical perspective using fossil charophytes. These beds have yielded a diverse charophyte assemblage, consisting of Platychara perlata, Platychara raoi, Platychara sahnii, Peckichara cf. varians, Nemegtichara cf. grambasti, and Chara sp. 1. These charophytes were recovered from a 60 cm thick clayey and nodular limestone unit that is underlain by claystone and overlain by laminated claystones deposited on fluvial floodplains. This charophyte assemblage suggests a Late Cretaceous–Early Danian age, which agrees with the previous dating carried out using freshwater to brackish water ostracods and planktic foraminifera found in the same beds, together with dasycladalean and halimedacean chlorophytes. The Jhilmili intertrappean charophyte assemblage resembles those found in other infra‐and intertrappean sections of western, central, and southern India. The charophyte assemblage indicates that during the Cretaceous–Palaeogene timespan the Indian subcontinent mainly experienced interchange with Laurasia (China, Europe, and North America) but also with Gondwana (North Africa and South America). In the opposite sense, Laurasian species might have dispersed to India by means of a sweepstakes route across the Kohistan–Dras volcanic arcs.

Ordovician stratigraphy and benthic community replacements in the eastern Anti-Atlas, Morocco

Abstract: Abstract The Anti-Atlas contains a thick, volcanic-free Ordovician succession that originally deposited in a passive-margin basin. Three main sedimentary packages are bounded by major unconformities: (i) the Tremadocian–Floian Lower Fezouata and Upper Fezouata formations, which unconformably overlie a palaeorelief of Cambrian rifting volcanosedimentary complexes, and are subsequently topped by a Dapingian paraconformable gap; (ii) the Darriwilian–Katian Tachilla Formation and First Bani and Ktaoua groups, the latter unconformably overlain by a Hirnantian glaciogenic succession; and (iii) the Second Bani Group, which subsequently infilled the former glaciogenic palaeorelief. Due to the scarcity of carbonate interbeds for etching analyses, leading to rare references of conodonts, the global Ordovician chart is interpolated on the basis of microphytoplancton (acritarchs and chitinozoans), regional graptolites and brachiopods. The Ordovician counter-clockwise rotation of Gondwana led its Moroccan margin from mid- to high-latitude positions, leading to the onset of a siliciclastic, wave- and storm-dominated platform. Flooding surfaces are marked by shelly silty carbonate interbeds that reflect the episodic development of echinoderm–bryozoan meadows during Katian times; in areas protected from siliciclastic input, they reached massive and bedded bioaccumulations (Khabt-el-Hajar Formation). The Hirnantian glaciation controlled the incision of numerous tunnel channels, infilled with both alluvial to fluvial sediments and glaciomarine diamictites. The Hirnantian palaeorelief was definitively sealed during Silurian times.

Paleo-Tethyan Ocean Evolution and Indosinian Orogenesis in the East Kunlun Orogen, Northern Tibetan Plateau

Abstract: The East Kunlun Orogen on the northern margin of the Tethyan orogenic system records a history of Gondwana dispersal and Laurasian accretion. Uncertainties remain regarding the detailed histories of northern branches of the Paleo-Tethys Ocean in East Kunlun Orogen (Buqingshan Ocean). Based on a synthesis of sedimentary, structural, lithological, geochemical, and geochronological data from the East Kunlun Orogen and adjacent regions, this paper discusses the spreading and northward consumption of the Paleo-Tethys Ocean during Late Paleozoic–Early Mesozoic times. The main evolutionary stages are: (1) during Carboniferous to Middle Permian, the Paleo-Tethys Ocean (Buqingshan Ocean) was in an ocean spreading stage, as suggested by the occurrence of Carboniferous MORB-, and OIB-type oceanic units and Carboniferous to Middle Permian Passive continental margin deposits; (2) the Buqingshan Ocean subducted northward beneath the East Kunlun Terrane, leading to the development of a large continental magmatic arc (Burhan Budai arc) and forearc basin between ~270–240 Ma; (3) during the late Middle Triassic to early Late Triassic (ca. 240–230 Ma), the Qiangtang terrane collided with the East Kunlun–Qaidam terranes, leading to the final closure of the Buqingshan Ocean and occurrences of minor collision-type magmatism and potentially inception of the Bayan Har foreland basin; (4) finally, the East Kunlun Orogen evolved into a post-collisional stage and produced major magmatic flare-ups and polymetallic mineral deposits between Late Triassic to Early Jurassic (ca. 230–200 Ma), which is possibly related to asthenospheric mantle upwelling induced by delamination of thickened continental lithosphere and partial melting of the lower crust. In this paper, we propose that the Wilson cycle-like processes controlled the Late Paleozoic–Early Triassic tectonic evolution of East Kunlun, which provides significant implications for the evolution of the Paleo-Tethys Ocean.

Relics of ophiolite-bearing accretionary wedges in NE Brazil and NW Africa: connecting threads of western Gondwana´s ocean during Neoproterozoic times

Abstract: Neoproterozoic breakup of Rodinia resulted in the formation of several oceanic realms between dispersing cratons, which were later consumed during the assembly of Gondwana. In its western portion, the interior orogenic belts of Gondwana formed during the Brasiliano-Pan African Orogeny in the late Neoproterozoic-early Cambrian. Available geophysical, structural and petrological data suggest that the complex network of shear zones that once connected the Borborema province (NE Brazil), Tuareg shield (Hoggar) and Central African domain (NW Africa) likely represent ancient sutures that mark collisional episodes between Archean-Paleoproterozoic paleocontinents such as Amazonian-West African and São Francisco-Congo. Mafic, ultramafic and sedimentary sequences associated with this set of structures respresent dismembered ophiolite slices interpreted as oceanic remnants (sensu lato) that were emplaced during the late stages of the Gondwana assembly. For instance, the composite Transbrasiliano-Khandi-In-Tedeini-Silet shear system crosscuts rock assemblages preserving a complex history of oceanic-crust-transition development (Novo Oriente complex) in association with primitive to evolved magmatic arcs and UHP rocks both in the Borborema province and NW Africa. In the central Borborema province, preserved ophiolitic slices are strongly overprinted by ductile and brittle deformation events, but partially preserved MORB-like amphibolites are akin to subduction-related-types that crystallized in early- and late Neoproterozoic times docked via terrane accretion and dispersed by strike-slip shear zones. In the southern Borborema province, an example of a Neoproterozoic ophiolitic assemblage is the Monte Orebe complex, that encompasses T-MORB mafic rocks, ultramafic lenses, and exhalative sedimentary rocks akin to early to late stages of oceanic basin spreading, emplaced during convergent plate motions between the Pernambuco-Alagoas superterrane and the São Francisco craton. Correlative units are found in Cameroon, including the strongly hydrotermalized ultramafic rocks of the Lomié and Boumnyebel complexes, that are structurally controlled by top-to-the-south verging nappes found in the N-NW margin of the Congo craton. In all scenarios, the ophiolitic complexes are related to intra-oceanic and continental magmatic arcs as well as to geophysical signatures comparable to Phanerozoic suture zones. Although strongly dismembered, scrapped off Neoproterozoic oceanic crust partially preserved within the major belts of western Gondwana demonstrate the role of accretion-collisional orogenesis during its assembly.

Paleoproterozoic emplacement and Cambrian ultrahigh-temperature metamorphism of a layered magmatic intrusion from the Central Madurai Block, southern India: From Columbia to Gondwana

Abstract: The Madurai Block in the Southern Granulite Terrane (SGT) of Peninsular India is one of the largest crustal blocks within the Neoproterozoic Gondwana assembly. This block is composed of three sub-blocks: the Neoarchean Northern Madurai block, Paleoproterozoic Central Madurai block and the dominantly Neoproterozoic Southern Madurai Block. The margins of these blocks are well-known for the occurrence of ultrahigh-temperature (UHT) granulite facies rocks mostly represented by Mg-Al metasediments. Here we report a dismembered layered mafic–ultramafic intrusion occurring in association with Mg-Al granulites from the classic locality of Ganguvarpatti in the Central Madurai Block. The major rock types of the layered intrusion include spinel orthopyroxenite, garnet-bearing gabbro, gabbro and gabbroic anorthosite showing rhythmic stratification and cumulate texture. The orthopyroxene-cordierite granulite from the associated Mg-Al layer is composed of spinel, cordierite and orthopyroxene. The pyroxene in both rock units is high-Al orthopyroxene formed under UHT metamorphic conditions. Conventional thermobarometry yields near-peak metamorphic conditions of 9.5–10 kbar pressure and a minimum temperature of 980 °C. We computed P–T pseudosections and contoured for the compositional as well as modal isopleths of the major mineral phases, which yield temperature above 1000 °C. FMAS petrogenetic grid, Al-in-orthopyroxene isopleth, conventional thermobarometry and calculated pseudosection reveal a clockwise pressure–temperature (P–T) path and near isothermal decompression. The U–Pb data on zircon grains from the layered magmatic suite indicate emplacement of the protolith at ca. 2.0 Ga and the metamorphic overgrowths yield weighted 206Pb/238U mean ages ca. 520 Ma. Monazite from the garnet-bearing gabbro and Opx-Crd granulite yielded 206Pb/238U weighted mean ages of ca. 532 Ma and 523 Ma marking the timing of metamorphism. We correlate the layered intrusion to a Paleoproterozoic suprasubduction zone setting, defining the Ganguvarpatti area as part of a collisional suture assembling the Northern and Central Madurai Blocks. The Paleoproterozoic magmatism and late Neoproterozoic-Cambrian UHT metamorphism can be linked to the tectonics of the Columbia and Gondwana supercontinents.

Paleoproterozoic emplacement and Cambrian ultrahigh-temperature metamorphism of a layered magmatic intrusion from the Central Madurai Block, southern India: From Columbia to Gondwana

Abstract: The Madurai Block in the Southern Granulite Terrane (SGT) of Peninsular India is one of the largest crustal blocks within the Neoproterozoic Gondwana assembly. This block is composed of three sub-blocks: the Neoarchean Northern Madurai block, Paleoproterozoic Central Madurai block and the dominantly Neoproterozoic Southern Madurai Block. The margins of these blocks are well-known for the occurrence of ultrahigh-temperature (UHT) granulite facies rocks mostly represented by Mg-Al metasediments. Here we report a dismembered layered mafic–ultramafic intrusion occurring in association with Mg-Al granulites from the classic locality of Ganguvarpatti in the Central Madurai Block. The major rock types of the layered intrusion include spinel orthopyroxenite, garnet-bearing gabbro, gabbro and gabbroic anorthosite showing rhythmic stratification and cumulate texture. The orthopyroxene-cordierite granulite from the associated Mg-Al layer is composed of spinel, cordierite and orthopyroxene. The pyroxene in both rock units is high-Al orthopyroxene formed under UHT metamorphic conditions. Conventional thermobarometry yields near-peak metamorphic conditions of 9.5–10 kbar pressure and a minimum temperature of 980 °C. We computed P–T pseudosections and contoured for the compositional as well as modal isopleths of the major mineral phases, which yield temperature above 1000 °C. FMAS petrogenetic grid, Al-in-orthopyroxene isopleth, conventional thermobarometry and calculated pseudosection reveal a clockwise pressure–temperature (P–T) path and near isothermal decompression. The U–Pb data on zircon grains from the layered magmatic suite indicate emplacement of the protolith at ca. 2.0 Ga and the metamorphic overgrowths yield weighted 206Pb/238U mean ages ca. 520 Ma. Monazite from the garnet-bearing gabbro and Opx-Crd granulite yielded 206Pb/238U weighted mean ages of ca. 532 Ma and 523 Ma marking the timing of metamorphism. We correlate the layered intrusion to a Paleoproterozoic suprasubduction zone setting, defining the Ganguvarpatti area as part of a collisional suture assembling the Northern and Central Madurai Blocks. The Paleoproterozoic magmatism and late Neoproterozoic-Cambrian UHT metamorphism can be linked to the tectonics of the Columbia and Gondwana supercontinents.

A new cynodont from the Upper Triassic Los Colorados Formation (Argentina, South America) reveals a novel paleobiogeographic context for mammalian ancestors

Abstract: Abstract Probainognathia is a derived lineage of cynodonts which encompass Mammalia as their crown-group. The rich record of probainognathians from the Carnian of Argentina contrasts with their Norian representation, with only one named species. Here we describe a new probainognathian, Tessellatia bonapartei gen. et sp. nov., from the Norian Los Colorados Formation of the Ischigualasto-Villa Unión Basin of Argentina. The new taxon, represented by a partial cranium with associated lower jaws, was analyzed through neutron and X-rays micro-tomography (μCT). The high-resolution neutron μCT data allowed the identification of a unique character combination, including features inaccessible through traditional techniques. We constructed the largest phylogenetic data matrix of non-mammalian cynodonts. The new species and its sister taxon, the Brazilian Therioherpeton cargnini , are recovered as probainognathians, closely related to Mammaliamorpha. We conducted the first quantitative paleobiogeographic analysis of non-mammalian cynodonts, focusing in probainognathians. The results indicate that Probainognathia and Mammaliamorpha originated in southwestern Gondwana (in the Brazilian Paraná Basin), which was an important center of diversification during the Triassic. Finally, the Chinese Lufeng Basin is identified as the ancestral area of Mammaliaformes. These new findings, besides adding to the knowledge of the poorly represented Norian cynodonts from the Los Colorados Formation, are significant to improve our understanding of probainognathian diversity, evolution, and paleobiogeographic history.

Three dispersal routes out of Africa: A puzzling biogeographical history in freshwater planarians

Abstract: Freshwater planarians may have a wide geographical range despite their assumed low vagility. Found across four continents, Dugesia may have either an ancient origin on a large palaeo landmass, followed by colonisation in different regions before continental fragmentation, or a more recent origin and subsequent transoceanic dispersal. We seek to resolve between these two hypotheses.

A large Megaraptoridae (Theropoda: Coelurosauria) from Upper Cretaceous (Maastrichtian) of Patagonia, Argentina

Abstract: Megaraptora is a theropod clade known from former Gondwana landmasses and Asia. Most members of the clade are known from the Early to Late Cretaceous (Barremian-Santonian), with Maastrichtian megaraptorans known only from isolated and poorly informative remains. The aim of the present contribution is to describe a partial skeleton of a megaraptorid from Maastrichtian beds in Santa Cruz Province, Argentina. This new specimen is the most informative megaraptoran known from Maastrichtian age, and is herein described as a new taxon. Phylogenetic analysis nested the new taxon together with other South American megaraptorans in a monophyletic clade, whereas Australian and Asian members constitute successive stem groups. South American forms differ from more basal megaraptorans in several anatomical features and in being much larger and more robustly built.