Showing papers in "Palaeobiodiversity and Palaeoenvironments in 2021"

The topographic evolution of the Tibetan Region as revealed by palaeontology

Abstract: The Tibetan Plateau was built through a succession of Gondwanan terranes colliding with Asia during the Mesozoic. These accretions produced a complex Paleogene topography of several predominantly east–west trending mountain ranges separated by deep valleys. Despite this piecemeal assembly and resultant complex relief, Tibet has traditionally been thought of as a coherent entity rising as one unit. This has led to the widely used phrase ‘the uplift of the Tibetan Plateau’, which is a false concept borne of simplistic modelling and confounds understanding the complex interactions between topography climate and biodiversity. Here, using the rich palaeontological record of the Tibetan region, we review what is known about the past topography of the Tibetan region using a combination of quantitative isotope and fossil palaeoaltimetric proxies, and present a new synthesis of the orography of Tibet throughout the Paleogene. We show why ‘the uplift of the Tibetan Plateau’ never occurred, and quantify a new pattern of topographic and landscape evolution that contributed to the development of today’s extraordinary Asian biodiversity.

The Devonian–Carboniferous boundary in the Carnic Alps (Austria and Italy)

Abstract: In the Carnic Alps there are four sections exposing rocks across the Devonian–Carboniferous boundary: the Grune Schneid and the Kronhofgraben sections in Austria, the Plan di Zermula A and the Sentiero per Cresta Verde sections in Italy. All of them are mainly composed of limestone and span from the late Famennian through the Tournaisian. In the Kronhofgraben and Plan di Zermula A sections, the limestone sedimentation is interrupted by black shales interpreted as equivalent of the Hangenberg Black Shales. The Grune Schneid and the Sentiero per Cresta Verde sections are two of the few sections worldwide, where the calcareous sedimentation is continuous. The main extinction event registered at the end of the Devonian is testified not only in the sections where the calcareous sedimentation was replaced by black shales sedimentation but also in the Grune Schneid and in the Sentiero per Cresta Verde sections. Data on conodont biostratigraphy and biofacies, the content in ammonoids, and trilobites and geochemistry patterns from all sections are here summarised and reviewed. These data-set allow to conclude that the four Carnic Alps sections positively test the suitability of the timeline proposed by the DCB working group for the definition of the revised Devonian–Carboniferous boundary.

The Devonian-Carboniferous boundary in the United States

Abstract: The Devonian-Carboniferous boundary is documented in marine strata from the Appalachian Basin, Michigan Basin, Illinois Basin, Forest City Basin, Williston Basin, Tobosa Basin, southern continental margin, western platform, and several sub-basins of the Antler Foreland in the USA based on conodonts, brachiopods, miospores, carbon isotope data, and magnetic susceptibility data. The boundary is narrowly constrained and conformable in the Illinois Basin and Oklahoma; less well constrained in the Appalachian Basin, Antler Foreland, western platform, and the Alberta Platform, and often developed on an unconformity.

Devonian-Carboniferous boundary sections in Iran

Abstract: Many sections are known from Iran which exhibit sediments across the Devonian-Carboniferous (D-C) boundary. In contrast to the majority of published D-C sections worldwide from pelagic/hemipelagic environments, successions in Iran are mainly composed of shallow-water sediments. Correlation with hemipelagic or pelagic palaeoenvironments remains difficult due to biostratigraphic uncertainties in most sections and/or hiatuses. On the other hand, a limited number of sections dealing with shallow-water facies settings in Iran at this particular time period are known and further research is necessary. Several sections in the Alborz Mountains provide an excellent opportunity to study successions across the D-C boundary in shallow-water facies. In Iran, protognathoids are represented by Protognathodus meischneri and Protognathodus collinsoni. The two biostratigraphically important protognathoids (Protognathodus kuehni and Protognathodus kockeli) were not reported or did not occur for the first time in the Late Tournaisian. Early siphonodellids were described instead. In the frame of an Iranian/German research project, we study different palaeoenvironments to reduce serious palaeoenvironmental and palaeogeographical sampling bias which may limit our knowledge on the Hangenberg Event particularly in shallow-water facies. We present a summary on published D-C sections in Iran (Ghale-Kalaghu, Howz-e-Dorah 1, Howz-e-Dorah 2 and Shahmirzad) and sections which are under study (Mighan, Chelcheli and Khoshyeilagh) at the time of this writing.

The Hushoot Shiveetiin gol section (Baruunhuurai Terrane, Mongolia): sedimentology and facies from a Late Devonian island arc setting

Abstract: A Late Devonian to (?)Early Mississippian section at Hushoot Shiveetiin gol in the Baruunhuurai Terrane of the Central Asian Orogenic Belt (CAOB) exposes large parts of cyclic Famennian shallow-water siliciclastic shelf deposits composed of siltstones, sandstones, shales, volcaniclastics, and intercalated autochthonous carbonates. The youngest part of the section, possibly Early Mississippian, is represented by arkosic sandstones with large plant remains. The facies reflects a range from shallow-intertidal to outer ramp settings. In terms of conodont stratigraphy, the Hushoot Shiveetiin gol section ranges from the Palmatolepis minuta minuta Biozone to at least the Palmatolepis rugosa trachytera Biozone. Hiatuses of several conodont biozones occur due to the facies setting (erosion and reworked sediments which are recognized by reworked conodonts) rather than thrusting or folding. The environmental setting was characterized by coeval subaerial volcanism resulting in numerous pyroclastic deposits. The depositional environments and intense volcanic activity at the Hushoot Shiveetiin gol section limited the stratigraphic distribution, abundance, and diversity of many elements of the fauna such as brachiopods. Ostracods were very abundant and diverse through many parts of the section. Although limited in stratigraphic distribution, the crinoid fauna is the most diverse Palaeozoic fauna collected from Mongolia to date and supports the hypothesis that the CAOB was a biodiversity hotspot in the aftermath of the Frasnian–Famennian extinction event.

Review of Devonian-Carboniferous Boundary sections in the Rhenish Slate Mountains (Germany)

Abstract: Thirty Devonian-Carboniferous Boundary sections of the Rhenish Slate Mountains and adjacent subsurface areas are reviewed with respect to litho-, event, conodont, ammonoid, sequence, and chemostratigraphy. In the interval from the base of the uppermost Famennian (Wocklum Beds, Wocklumian) to the base of the middle Tournaisian (base Lower Alum Shale), 11 conodont and 16 ammonoid (sub)zones are distinguished. The terminology of the Hangenberg Crisis Interval is refined, with an overall regressive Crisis Prelude below the main Hangenberg Extinction, which defines the base of the transgressive Lower Crisis Interval (Hangenberg Black Shale). The glacigenic and regressive Middle Crisis Interval (Hangenberg Shale/Sandstone) is followed by the overall transgressive Upper Crisis Interval that can be subdivided into three parts (I to III) with the help of conodont stratigraphy (upper costatus-kockeli Interregnum = upper ckI, Protognathodus kockeli Zone, and lower part of Siphonodella (Eosiphonodella) sulcata s.l./Pr. kuehni Zone). Protognathodus kockeli includes currently a wide range of forms, which variabilities and precise ranges need to be established before a precise GSSP level should be selected. Returning to its original definition, the former Upper duplicata Zone is re-named as Siphonodella (S.) mehli Zone. It replaces the S. (S.) jii Zone, which is hampered by taxonomic complications. The S. (S.) quadruplicata Zone of Ji (1985) is hardly supported by Rhenish data. The entry of typical S. (S.) lobata (M1) characterises an upper subdivision (subzone) of the S. (S.) sandbergi Zone; the new S. (S.) lobata M2 enters much earlier within the S. (S.) mehli Zone. The ammonoid-defined base of the Wocklum-Stufe (Upper Devonian = UD VI) begins with the Linguaclymenia similis Zone (UD VI-A1). The oldest S. (Eosiphonodella) enter within the Muessenbiaergia bisulcata Zone (UD VI-A2). The traditional Parawocklumeria paradoxa Zone of Schindewolf (1937) is divided into successive P. paprothae (VI-C1), P. paradoxa (VI-C2), and Mayneoceras nucleus (VI-C3) Subzones. In the lower Tournaisian (Lower Carboniferous = LC I), the Gattendorfia subinvoluta Zone is subdivided into G. subinvoluta (LC I-A2) and “Eocanites” nodosus (LC I-A3) Subzones. The Paprothites dorsoplanus Zone (LC I-B) can be divided into Pap. dorsoplanus (LC I-B1) and Paragattendorfia sphaeroides (LC I-B2) Subzones. Potential subdivisions of the Pseudarietites westfalicus (LC I-C) and Parag. patens Zones (LC I-D) are less distinctive. The unfossiliferous or argillaceous upper part of the Hangenberg Limestone and the overlying Lower Alum Shale Event Interval remain regionally unzoned for ammonoids.

The Devonian–Carboniferous boundary in the stratotype area (SE Montagne Noire, France)

Abstract: Sections with continuous sedimentation across the Devonian–Carboniferous (D-C) boundary in the Montagne Noire allow to build a virtual transect from shoreline to deep basin. Nearshore facies characterise the D-C boundary stratotype and neighbouring sections at La Serre in the Cabrieres klippen domain, and offshore facies are present at the Col de Tribes and Puech de la Suque sections in the Mont Peyroux nappe domain. Both domains exhibit equivalents of the Hangenberg Black Shale (HBS). At La Serre, an initial regressive trend is indicated by the presence of oculated trilobites in the topmost pre-HBS Wocklumeria Limestones. Above the HBS level, regressive depositional conditions characterise oolitic deposits that comprise lithic erosional flows with an admixture of transported shallow-water biotas. Maximum regression is recognised with the deposition of coarse breccias and local features of emergence prior to the first appearance of Protognathodus kockeli. The oolites are superseded by the transgression of outer shelf deposits. In the nappe domain, the HBS is intercalated in outer ramp nodular limestones, and it exhibits detrital elements pointing to its regressive nature. The regressive trend culminates than reverses when post-HBS carbonate sedimentation resumes. Protognathodus kockeli appears in the post-HBS carbonates. Associated oculated trilobites indicate shallower bathymetric conditions then those of the pre-HBS Wocklumeria Limestones. Thereafter, replacement of sighted trilobites by blind ones and the protognathodid biofacies by facies dominated by siphonodellids indicate a deepening trend. The near- and offshore sites of the D-C transition permit correlation of short-term bathymetric fluctuations with faunal turnovers and entries of biostratigraphic markers.

The Devonian/Carboniferous boundary interval in Poland: multidisciplinary studies in pelagic (Holy Cross Mountains and Sudetes) and ramp (Western Pomerania) successions

Abstract: A multidisciplinary study of the Devonian–Carboniferous boundary interval in pelagic successions of the Holy Cross Mountains and Sudetes and the ramp successions in the Western Pomerania region (Poland) is presented herein. The analysis applies the results of new palaeontological and biostratigraphic studies based mainly on conodonts, ammonoids and palynomorphs, biostratigraphic results interpreted earlier by different authors that have been re-examined, and geochemical and mineralogical characteristics, as well as magnetic susceptibility measurements across the Devonian–Carboniferous boundary interval. The study is focused on the interval from the Famennian ultimus conodont Zone to the Tournaisian duplicata conodont Zone, and from the Famennian lepidophyta–explanatus (LE) miospore Zone to the Tournaisian verrucosus–incohatus (VI) miospore Zone, respectively. The paper highlights sections, which are the most representative for the Devonian–Carboniferous boundary in each region, illustrates and summarises current knowledge on the uppermost Famennian to lowermost Tournaisian in these regions, gives data and correlation of the important stratigraphic markers for each region, and briefly correlates them outside the region. The sedimentary successions and specific phenomena, together with microscale environmental perturbations, recognised close to the Devonian/Carboniferous boundary in Poland, display a pattern similar to that observed in many areas in Europe during the Hangenberg Event.

Devonian-Carboniferous boundary in China

Abstract: The Devonian-Carboniferous transition is marked by major perturbations to the Earth’s climate system and marine redox conditions, a first-order mass extinction, and the succeeding marine ecosystem recovery, recording a sequence of environmental and biotic events. In South China, there exist numerous well-preserved and continuous Devonian-Carboniferous boundary successions that were formed in low-latitude passive margin basin, and this presents a unique opportunity to decipher the strata along a proximal to basinal transect using integrated biostratigraphical, event-stratigraphical, and geochemical approaches. Our geochemical data (δ13Ccarb, δ15Nbulk, δ238U and Mo/U enrichments, I/Ca ratio) indicated local and global C-N cycling perturbations and marine anoxia in the main phase of the Hangenberg Crisis interval (Middle Siphonodella praesulcata Zone), which coincided with a major regression in South China. Stromatoporoid biostromes and most typical Devonian faunas did not survive into the interval, and the deep-water black shales yield only opportunistic survivors, such as ammonoid Postclymenia cf. evoluta and bivalve Guerichina, along with miospores belonging to the LN Zone. The upper Hangenberg Crisis interval (Upper Si. praesulcata Zone, equal to the Protognathodus kockeli Zone) is marked by initial post-glacial transgression, a global δ13Ccarb spike, and opportunistic faunal blooms, coincident with conodont biofacies shift to the polygnathid-protognathodid biofacies. The Protognathodus fauna, nearly absent from platform facies, is often found in the condensed upper crisis interval of the basin, slope, and platform margin facies within the Youjiang Basin, although the abundance is extremely low and the phylogenetic lineage of Pr. meischneri-Pr. collinsoni-Pr. kockeli cannot be recognised in most successions. In South China, the first occurrence of Pr. kockeli seems to correspond to lithofacies changes, recorded in the first level of transgression just above the Hangenberg Sandstone event. Multiple lines of evidence, including great spatial heterogeneity of δ13Ccarb and δ15Nbulk records, extremes of redox proxies, and lack of macrofossils, suggest that marine habitats remained in a critical/turbulent state during the Upper Si. praesulcata Zone, triggering the opportunistic blooms and minor extinctions. Following the crisis intervals, radiation/recovery of many fossil groups did not occur until the upper part of the Siphonodella sulcata Zone to the Si. duplicata Zone. In particular, a negative shift of δ13Ccarb to pre-crisis values occurring near the base of the Si. sulcata/Pr. kuehni Zone could be recognised worldwide. The research history, current status, and achievement of Devonian-Carboniferous boundary in China are briefly reviewed.

A terrestrial Devonian-Carboniferous boundary section in East Greenland

Abstract: Terrestrial Devonian-Carboniferous boundary sections are present in the East Greenland Devonian Basin. The boundary section on Stensio Bjerg developed in deep, distal lake sediments with a pair of lakes representing the boundary. A diverse spore assemblage developed as the lake flooded the basin. Previously abundant spores, notably Retispora lepidophyta, Diducites spp., Rugospora radiata and all forms with bifurcate tips (Ancyrospora and Hystricosporites), then became extinct through just over a metre of section. The spore assemblage is then lost into AOM rich very high TOC% lake sediments. There is a negative δ13CTOC excursion in the Stensio Bjerg section interpreted to represent the upper part of the positive excursion known from marine sections. The upper lake contains the simple VI spore assemblage of the earliest Carboniferous age. The correlative section on Rebild Bakker was developed in shallow proximal facies without AOM and shows that a Devonian-Carboniferous LN* to VI spore zone boundary can be picked in the lower lake based on the last occurrence of Retispora lepidophyta in an assemblage otherwise dominated by simple spores and Grandispora cornuta. Spores in this VI spore assemblage, particularly Grandispora cornuta, show sculpture malformation that is entirely characteristic of UV-B radiation damage to their DNA prior to deposition of its protective wall layer. This palynological record showing the rapid extinction of major elements within the Late Devonian microflora can be reconciled with accounts claiming there was no mass extinction of plants and spores across the boundary. The palaeobiology of the major spore groups that became extinct is reviewed.

Late Devonian (Famennian) phacopid trilobites from western Mongolia

Abstract: The early Famennian phacopid trilobites Feistops mongoliensis gen. nov. sp. nov., Feistops khovdensis gen. nov. sp. nov., ?Feistops sp. A, and ?Houseops olonbulagensis sp. nov. are recorded and illustrated from the western Mongolia Hushoot Shiveetiin gol section from the Baruunhuurai Terrane of the Central Asian Orogenic Belt (CAOB). In this section, exposing large parts of cyclic Famennian shallow-water siliciclastic shelf deposits, Feistops mongoliensis gen. nov. sp. nov., Feistops khovdensis gen. nov. sp. nov., ?Feistops sp. A, and ?Houseops olonbulagensis sp. nov. are the first known and described representatives of trilobites from Late Devonian deposits of western Mongolia. Their affinities with known other sighted species of the same age are discussed.

The Devonian/Carboniferous boundary in Sardinia (Italy)

Abstract: In Sardinia (Italy) two sections, located in the southeastern part of the island, expose the Devonian/Carboniferous Boundary: the Monte Taccu Section and the Bruncu Bullai section. Both expose limestones from the middle Famennian to the lower Tournaisian Siphonodella jii Zone. The limestone sequence is interrupted by a thin level of shales representing the Hangenberg Event. Data on conodont biostratigraphy and magnetic susceptibility across the Devonian/Carboniferous Boundary are here summarised and reviewed.

The Devonian–Carboniferous boundary in Belgium and surrounding areas

Abstract: The Devonian–Carboniferous boundary is associated with a major extinction event of the Phanerozoic. It was also a time marked by a rapid but short-lasting change in deposition called Hangenberg Event. In the Namur–Dinant Basin the uppermost Devonian (‘Strunian’) deposits recorded a third-order transgression that produced a progressive switch from coastal siliciclastic to proximal mixed deposits with an increase of the carbonate production on the ramp. Hence, the Comblain-au-Pont and lower Hastiere formations are considered as the transgressive system tract, whereas the middle member of the Hastiere Formation is interpreted as the highstand system tract, capped by an erosion surface corresponding to the third-order sequence boundary. Superimposed on these third-order sequences are well-marked orbitally forced precession cycles (wet–dry climate alternations) of c. 18.6 ka, appearing as irregular c. 30–80-cm-thick couplets of limestone and calcareous shale beds. The Hangenberg Black Shale Event is locally present as dark shales that likely spread over the shelf, marking the maximum flooding surface of the sequence. Before and after this event, carbonate facies rich in benthic macrofauna and microfauna continued to develop. The Hangenberg Sandstone Event, appearing as a sandstone bed in pelagic sections, is variously recorded at the base of the Hastiere Formation, either as a sandy siltstone bed in proximal sections or as a horizon with limestone clasts and reworked fossils in more distal settings. The Hangenberg Sandstone Event beds occur sharply in the stratigraphic record and do not correspond to the long sea level fall of a third-order sequence boundary, but most probably to a short out-of-sequence event. The revision of the stratigraphic distribution of major fossil groups pleads for a continuous biostratigraphic succession with no obvious hiatus. The variable development of some micropalaeontological zones at the end of the Devonian is the result of complex eco-biostratigraphic interactions with the environment rather than the reflection of true hiatuses. It is marked by extinctions of Devonian taxa, concomitantly with the end of the reworking produced by the Hangenberg Sandstone Event, most probably immediately below the entry of the conodont Protognathodus kockeli. It is also coincident with the boundary between the foraminiferal zones DFZ7–MFZ1, rugose coral zones RC0–RC1 and between the palynozones LE–VI. After the short-lasting regressive phase of the Hangenberg Sandstone Event, normal depositional settings returned with the deposition of the Hastiere Formation. Hence, the end of the Hangenberg Sandstone Event is proposed as the most natural proxy to pinpoint the Devonian–Carboniferous boundary.

The foraminiferal zonal scale of the Devonian–Carboniferous boundary beds in Russia and Western Kazakhstan and its correlation with ammonoid and conodont scales

Abstract: This paper reviews the foraminiferal biostratigraphy of the Devonian–Carboniferous (D–C) boundary beds in Russia and Western Kazakhstan. The regional records of the zonal successions of Russia are summarised from sections and boreholes of the Volga-Urals Subregion of the East European Platform (Syzran no. 401 Borehole, Melekesskaya no.1 Borehole); Timan-Pechora Province of the East European Platform (Kamenka Section); South Urals (Sikaza, Zigan, Ryauzyak and Dzerzhinka sections); Middle Urals (Pershino Section); and West Siberia, Vagai-Ishim Basin (Borehole no. 1 Kurgan-Uspenskaya), and the zonal successions of Western Kazakhstan are summarised from the Mugodzhary Mountains. Three foraminiferal biozones can be defined in the interval comprising the latest Devonian and earliest Carboniferous: Quasiendothyra kobeitusana Zone, the Tournayellina pseudobeata – remnant Quasiendothyra Zone, and the Earlandia minima Zone. The D–C boundary interval is situated within the Tournayellina pseudobeata–remnant Quasiendothyra Zone. A correlation with the ammonoid and conodont zonations is proposed, which is useful for correlating deep-water and shallow-water successions.

The impacts of diagenesis on the geochemical characteristics and Color Alteration Index of conodonts

Abstract: The conodont Color Alteration Index (CAI) has been widely used to determine the maximum temperature in carbonate rocks, despite recognition that conodont colour can be affected by other factors, such as diagenesis. Measurements of trace element characteristics in conodonts of varying CAI (1.5–6.0) from the Canadian Cordillera show that those specimens with the most anomalous CAI with respect to independent estimates of maximum temperature also have the highest concentrations of iron (> 3000 ppm). The adsorption of transition metals such as iron onto bioapatite crystals and permineralization by their oxides in conodont elements are herein proposed as mechanisms for the modification of CAI during diagenesis. Furthermore, the trace element characteristics of conodonts, primarily those of the lanthanide, or rare earth element (REE) series, have frequently been used as a proxy for palaeoceanographic conditions, including anoxic events. However, the impact of early diagenetic processes post-burial obscures this marine signal, and instead, the trace element characteristics of conodonts likely reflect the characteristics of pore waters or diagenetic fluids. Several geochemical ‘tools’ have been proposed to test for such overprinting of palaeoceanographic information, including Y/Ho vs. ΣREE, MREE/MREE*, U concentration, and La/Yb. However, REE and Raman structural characteristics in a suite of conodonts from the Canadian Cordillera indicate that these ‘tools’ cannot be systematically applied. The majority of conodont specimens analysed in this study appear to have been affected by post-burial diagenetic alteration using one or more of the ‘tools’, but seem to be unaltered when investigated using other ‘tools’. Additionally, several specimens which appear to be diagenetically unaltered when using the metrics of the geochemical ‘toolbox’ exhibit diagenetically induced structural changes. The geochemical ‘tools’ are thus unable to discriminate between diagenetic alteration induced before and after burial, and they should therefore be accompanied by structural analyses if REE characteristic of conodonts are used to infer palaeoceanographic conditions.

Famennian crinoids and blastoids (Echinodermata) from Mongolia

Abstract: Herein we report on the most abundant and diverse fauna of Palaeozoic crinoids and blastoids collected from Mongolia to date. The fauna is from the Late Devonian (Famennian) Samnuuruul Formation in western Mongolia. The fauna consists of two genera of blastoids and twelve genera of crinoids—four genera of camerates, three genera of flexibles, one disparid genus, and four genera of cladids. The crinoids and blastoids were living on an active island arc complex in the Central Asian Orogenic Belt (CAOB) in a high physical stress environment with frequent and often voluminous pyroclastic eruptions. The Mongolian fauna is similar to coeval faunas collected from the Hongguleleng Formation in western China and supports the hypothesis that the CAOB was a biodiversity hotspot for Famennian echinoderms and a precursor to the very successful echinoderm communities that dominated Mississippian shallow-marine ecosystems globally. Three new taxa are described. Mongoliacrinus minjini, new genus and species, is the oldest member of the Acrocrinidae, previously known from the Mississippian and Pennsylvanian and the first occurrence of the family outside North America. Eutaxocrinus ariunai and Eutaxocrinus sersmaai are new species of the flexible crinoid Eutaxocrinus, a genus with a widespread distribution during the Early and Middle Devonian, which survived into the Lower Mississippian. It is restricted to the CAOB in the Late Devonian.

Late Devonian ostracods (Crustacea) from the Hushoot Shiveetiin gol section (Baruunhuurai Terrane, Mongolia) and their palaeoenvironmental implication and palaeobiogeographic relationship

Abstract: An Upper Devonian to (?)Lower Mississippian section at Hushoot Shiveetiin gol in the Baruunhuurai Terrane of the Central Asian Orogenic Belt (CAOB) was investigated. A generally well-preserved and diverse ostracod fauna was collected from the Late Devonian (Famennian) Samnuuruul Formation in western Mongolia. It is the first rich assemblage described from this area. The ostracod fauna consists of 19 genera and 25 species. Two of the species are proposed new species, and 13 species are described in open nomenclature. The Mongolian ostracod fauna is similar to coeval faunas in western China and Laurussia, but also contains new, endemic species. Two new taxa are described: Beyrichiopsis hushootensis and Ampuloides beckeri. Ostracod findings are characterized by the Eifelian Mega-Assemblage (I-III) representing a nearshore, variable palaeoenvironment which is in accordance with facies analysis provided by other studies.

Linking modern-day relicts to a Miocene mangrove community of western Amazonia

Abstract: The Amazon drainage basin is extremely biodiverse, yet the origins of this diversification remain much debated. One of the possible drivers of plant diversity are the marine incursions that reached Amazonia during the Miocene and connected western Amazonia with the Caribbean. In the Miocene, large parts of western Amazonia were covered by extensive wetlands that, during high eustatic episodes, were episodically colonised by coastal taxa such as mangroves. In this paper, we hypothesise that some of these mangrove community taxa could adapt to freshwater conditions enriching the modern plant composition of the Amazon drainage basin. To assess the past plant composition in the basin, we statistically analyse the palynological composition of two Miocene sections from western Amazonia that were especially rich in presumed mangrove pollen (Zonocostites ramonae). We identify the pollen taxa that coexisted with this community using clustering methods supported by Kendall’s W coefficient concordance analysis. Our results suggest that at least fourteen taxa are closely associated with Zonocostites ramonae (Cricotriporites guianensis, Deltoidospora adriennis, Psilabrevitricolporites devriesii, Psiladiporites redundantis, Psilamonocolpites amazonicus, P. rinconii, Psilatricolporites crassoexinatus, P. labiatus, P. operculatus, Psilatriporites corstanjei, Retitricolporites kaarsii, Rhoipites guianensis, Rhoipites hispidus, and Tetracolporopollenites transversalis). We also illustrate the pollen of this fossil mangrove, and some of its associated fossil taxa, using light microscopy (LM) and scanning electron microscopy (SEM), and we compare them with potential nearest living relatives (NLR). We found that pollen of the modern mangrove Rhizophora mangle is the NLR of Zonocostites ramonae. Of the three associated taxa, the best analogy is between Psilabrevitricolporites devriesii and Humiria balsamifera, the latter a taxon best known from the coastal restinga vegetation in Brazil and Surinam. Tentatively, we assign Forsteronia spp. as NLR for Cricotriporites guianensis, and we propose Euterpe sp. for Psilamonocolpites rinconii (but also Oenocarpus, Hyospathe, Prestoea, and Sabinaria are affine). Based on this study we propose that, at least for some fossil taxa of the Miocene mangrove environment, there are still NLR or relict species that occur in inland Amazonia and along the South American coastline. We thus conclude that the Amazonian flora, like riverine fauna such as the pink river dolphin (Inia) and selected fish taxa, carry an imprint of the Miocene coastal communities.

The Devonian-Carboniferous Boundary around the globe: a complement

Abstract: This article compiles data on the Devonian-Carboniferous Boundary successions from countries and regions, which have not been dealt with separately in this special issue. Data derive from different palaeocontinents on a large palaeoclimatic gradient from the southern high latitudes of western Gondwana through the palaeotropics into the temperate/boreal northern latitudes of Siberia. The quality of the data is variable, but often surprisingly good, although major uncertainties can persist and traditional positions of the boundary do not withhold modern stratigraphic results and concepts. Not all these regions have been on the forefront of the DCB discussions in recent years, but they provide invaluable insights into the regional and global dimensions of uppermost Devonian and lowermost Carboniferous changes in the biosphere and geosphere. Almost globally documented are faunal turnovers and collapse of ecosystems in the Hangenberg extinctions. Those allow the placement of the DCB even if index fossils are absent or scarce. However, it also shows the regional differences, which advocate for a holistic approach to the boundary by ensuring global correlation without relying on the conceptual dominance a single taxon, which might be less powerful outside its usual facies.

Wildfire during deposition of the “Illinger Flözzone” (Heusweiler-Formation, “Stephanian B”, Kasimovian–Ghzelian) in the Saar-Nahe Basin (SW-Germany)

Abstract: Wildfires occurred more or less regularly in many Pennsylvanian ecosystems, not only in seasonally dry regions but also in the ever wet tropics. One of the reasons for this was probably the relatively high atmospheric oxygen conditions prevailing during this period. The present study reports evidence for the occurrence of wildfires during deposition of the Upper Pennsylvanian Heusweiler Formation (“Stephanian B”, Kasimovian–Gzhelian) in the intramontane Saar-Nahe (or Saar-Lorraine) Basin in SW-Germany. Based on anatomical features of the charcoal, as well as the co-occurring adpression flora, it seems possible that some of the fires occurred in an ecosystem inhabited by Cordaites. Some of the charcoal fragments exhibit traces of pre-charring decay by fungi, indicating either the consumption of litter by ground or surface fires, or of still standing (partly) dead trees by crown fires.

The stratigraphy of latest Devonian and earliest Carboniferous rocks in Ireland

Abstract: Thick successions of latest Devonian and early Carboniferous siliciclastic rocks of fluviatile and shallow marine origin are well exposed in numerous coastal sections in the south of Ireland. Correlations based on palynostratigraphy demonstrate that an overall northward transgression, in detail consisting of several fluctuations of relative sea level, resulted in the replacement of fluvial coastal plain environments by shallow marine conditions. The most conspicuous lithological record of transgression occurred at or close to the base of the Carboniferous, when a regionally distributed mudstone deposited in a shallow sea, replaced the generally sandy strata that had prevailed in the latest Devonian. There is no evidence for very large glacioeustatic changes of sea level related to the coeval western Gondwanan glaciation. Of the three horizons for a new GSSP for the base of the Carboniferous currently under review, the base of the Protognathodus kuehni Zone/basal Siphonodella sulcata Zone and coastal plant extinction is the most easily applied in Ireland and can be identified using palynology and corresponds to an easily recognised, extensively developed lithological change in the south of the country.

Late Pliocene to early Pleistocene climate dynamics in western North America based on a new pollen record from paleo-Lake Idaho

Abstract: Marked by the expansion of ice sheets in the high latitudes, the intensification of Northern Hemisphere glaciation across the Plio/Pleistocene transition at ~ 2.7 Ma represents a critical interval of late Neogene climate evolution. To date, the characteristics of climate change in North America during that time and its imprint on vegetation has remained poorly constrained because of the lack of continuous, highly resolved terrestrial records. We here assess the vegetation dynamics in northwestern North America during the late Pliocene and early Pleistocene (c. 2.8–2.4 Ma) based on a pollen record from a lacustrine sequence from paleo-Lake Idaho, western Snake River Plain (USA) that has been retrieved within the framework of an International Continental Drilling Program (ICDP) coring campaign. Our data indicate a sensitive response of forest ecosystems to glacial/interglacial variability paced by orbital obliquity across the study interval, and also highlight a distinct expansion of steppic elements that likely occurs during the first strong glacial of the Pleistocene, i.e. Marine Isotope Stage 100. The pollen data document a major forest biome change at ~ 2.6 Ma that is marked by the replacement of conifer-dominated forests by open mixed forests. Quantitative pollen-based climate estimates suggest that this forest reorganisation was associated with an increase in precipitation from the late Pliocene to the early Pleistocene. We attribute this shift to an enhanced moisture transport from the subarctic Pacific Ocean to North America, confirming the hypothesis that ocean-circulation changes were instrumental in the intensification of Northern Hemisphere glaciation.

Palynofacies as a palaeoenvironment and hydrocarbon source potential assessment tool: An example from the Cretaceous of north Western Desert, Egypt

Abstract: Optical examination employing transmitted light and UV-fluorescence microscopy of palynological preparations of eighteen cutting samples representing the Alam El Bueib Member (Hautervian-Barremian), Kharita/lower Bahariya (Cenomanian), and Abu Roash (Turonian-Santonian) formations collected from the Faghur Hj5-1 well, north Western Desert, Egypt, allows the identification of three different palynological assemblages from the studied rock units. These assemblages are mainly non-marine but apparently marine at the base of the Alam El Bueib Member, as evidenced by dinocyst occurrence. In addition, the presence of the Pediastrum and chlorophycean algae ecozone, recognised in previous works, is a good datum for the Abu Roash Formation in the north Western Desert of Egypt. Three associations of palynofacies linked to lithofacies changes are recognised and employed in identification of depositional environments. The Alam El Bueib samples yielded mixed kerogen assemblages of non-marine and marine organic facies. The Kharita/lower Bahariya interval is mostly barren, possibly due to prevailing sandstone lithofacies, except for one sample at its upper part which contains a diverse palynological assemblage. The overlying Abu Roash Formation has a homogeneous kerogen composition comprising mainly granular fluorescent AOM and algae as well as rare palynomorphs. Qualitative as well as quantitative variations of palynofacies allow the reconstruction of the depositional environment. The obtained data have the potential for discriminating spatial and redox status differences and providing also information about terrestrial/freshwater influxes. Results support the model that the Alam El Bueib Member was deposited in a marginal dysoxic-anoxic to distal suboxic-anoxic basin. The Kharita/lower Bahariya unit in the studied well was deposited under marginal dysoxic-anoxic conditions whereas the overlying Abu Roash Formation in a distal suboxic-anoxic basin. Palynofacies results also show that the studied material comprises two distinct facies of kerogen. First, Type II > I kerogen (AOM-rich) is overwhelmingly dominant in the Abu Roash Formation and a few samples from the Alam El Bueib Member which are presumed highly oil-prone, whereas Type III kerogen (phytoclast-rich) is particularly common in the Alam El Bueib Member and Kharita/lower Bahariya unit which are considered gas-prone. Thermal maturity determinations obtained from colour changes of smooth-walled palynomorphs reveal that Alam El Bueib samples belong to immature to mature stages; however, Kharita/lower Bahariya and Abu Roash samples are within the immature phase.

The anthracotheres from northern Junggar Basin and their palaeoclimatic significance in relation to the Tibetan Plateau

Abstract: Anthracotheres were an environmentally sensitive group that were thought to have become extinct from the middle-high latitudes of the northern hemisphere by the early Miocene. Hereby, the present author reports previously unpublished fossil bothriodonts, a group of anthracotheres, from the northern Junggar Basin, in China’s far northwest. An M3 of a small bothriodont Elomeryx sp., from the lower part of the Halamagai Formation, which is probably of the early middle Miocene age, is the only anthracothere record from the Neogene of northern China. More importantly, it is the latest anthracothere record from the middle-high latitudes of the northern hemisphere. The other teeth from Xiaerhete locality of the Irtysh River Formation, which is of the late Eocene age, are identified as Bakalovia sp. This poorly known genus was previously only reported from the middle or late Eocene of Southwest Europe. This study complements the sparse anthracothere fossil records of northwestern China. The present material, together with previous records, strongly suggests the occurrence of anthracothere interchanges between East Asia and both Europe and North America possibly from the middle or late Eocene to the early Oligocene. Anthracotheres declined remarkably in the Oligocene and Miocene in northern China, in contrast to their continued abundance in South and Southeast Asia at the same time, suggesting the existence of climate zonation and a faunal discontinuity in mammal distribution patterns between the northern and southern China. This zonation was probably related to the uplift of the Tibetan Plateau, which altered atmospheric circulation throughout Eastern Asia.

The Devonian-Carboniferous boundary in the Moravian Karst (Czech Republic)

Abstract: The continuous and well-accessible Devonian-Carboniferous boundary successions of the Czech Republic are part of the Brunovistulian Unit, which belongs to the Variscan Rhenohercynian Zone of the Bohemian Massif. The well-studied sections, with almost continuous carbonate sedimentation, crop out in the Moravian Karst area and are characterised by carbonate turbidites (Lesni lom) and hemipelagic successions (Křtiny). In the Lesni lom quarry, a complete succession of conodont zones of the Devonian-Carboniferous boundary interval (Upper expansa, praesulcata, costatus-kockeli, kockeli, sulcata/kuehni, bransoni, duplicata zones) was documented in calciturbidites and can be correlated with coeval foraminiferal zones (Quasiendothyra kobeitusana, Quasiendothyra konensis and Tournayellina pseudobeata zones). The Hangenberg Crisis and its three phases can be well recognised both with lithological, biostratigraphical and geochemical approaches in the Lesni lom section. At Křtiny, the nodular limestones yielded conodonts of the Devonian-Carboniferous boundary interval, but missing protognathodid fauna hampered identification of the kockeli Zone. A stratigraphic gap covering the basal part of the sulcata/kuehni Zone or strong condensation can be inferred from the occurrence of the transitional morphotype Si. sulcata-bransoni at the base of this zone. The tested “Montpellier criterion” for the Devonian-Carboniferous boundary redefinition can be used only in the Lesni lom section, where the first Protognathodus kockeli occurs above the regressive Hangenberg Sandstone Event deposits.

Pteridophytes as primary colonisers after catastrophic events through geological time and in recent history

Abstract: Pteridophytes reproduce by producing vast numbers of spores that may be dispersed over considerable distances, helping the plants colonise new areas. Being resistant to desiccation, fern spores can often survive for many years as spore banks in soil. After disturbance, such spores can germinate and subsequently colonise the area. These factors help pteridophytes to become primary colonisers on barren land, such as volcanic islands or land that has been devastated by some cataclysmic event. A further method of rapid colonisation is provided through the preservation and possible scattering of fragments of rhizomes in particular of horsetails. Similar rapid colonising by pteridophytes has been documented in the geological record following several major extinction events. These distinct, but short-lived, fern populations are recognisable by fern spikes in the microfossils. This paper brings together information on the reasons for pteridophyte success in colonising barren land, and examples taken from both the historic and geological records.

Compressions of Sequoia (Cupressaceae sensu lato) from the Middle Jurassic of Daohugou, Ningcheng, Inner Mongolia, China

Abstract: The specimens (two pairs of part and counterpart) of coniferalean branched leafy shoots were collected from the Jiulongshan Formation (Middle Jurassic) at the village of Daohugou, Ningcheng, Inner Mongolia, China. The fossil material displays helical phyllotaxy, dorsiventrally flattened linear, sessile leaves with decurrent base and prominent and single mid-vein. The foliate shoots are assigned to Sequoia jeholensis by comparing those of the genera Yanliaoa, Sequoia, Metasequoia, Taxodium, and Glyptostrobus, the basal members of the Cupressaceae sensu lato. Sequoia jeholensis is the oldest Sequoia species to date. It has coriaceous, dimorphic, helically arranged linear and scale leaves, with entire margins, attached obliquely to the axis by their decurrent bases. The leaf cuticle of S. jeholensis is described here for the first time. Leaves are hypostomatic, with stomata only on the abaxial surface. Anticlinal walls are straight, with single or more often paired pits. The epidermal structures of Sequoia are one of the key characters to distinguish it from similar genera, but they usually could not be used to distinguish the species within Sequoia. The Middle Jurassic of Daohugou represents a lacustrine environment with a warm and humid climate reflected by the species assemblage. So, the extinct Sequoia probably had similar climatic requirements to those of extant Sequoia sempervirens.

Climate and vegetation change during the Upper Siwalik—a study based on the palaeobotanical record of the eastern Himalaya

Abstract: Today, Northeastern India receives some of the highest annual rainfall totals globally. The major portion of annual precipitation in this region falls during the Indian Summer Monsoon season (June–September); however, this region also receives a significant amount of rainfall during the pre-monsoon season (March–May). Here, we quantitatively reconstruct the climate of the Upper Siwalik subgroup, eastern Himalayan region, based on fossil assemblages using the Coexistence Approach (CA). The age of the fossil assemblage is considered to be late Pliocene–early Pleistocene. Data reconstructed for the present-day and past pre-monsoon rainfall in Northeastern India indicate an increasing trend since the late Miocene–early Pliocene. During the late Pliocene–early Pleistocene (Upper Siwalik), the temperature seasonality between warm (27–28.1 °C) and cold months (22–23.6 °C) was less pronounced compared with present-day warm (27–27.7 °C) and cold (14.8–15.4 °C) months conditions at the fossil locality. The reconstructed rainfall data indicate a monsoonal type of climate having a strong seasonality in wet and dry seasons during the deposition of the Upper Siwalik sediments. Moreover, composition of the fossil floras and reconstructed palaeoclimate suggest a vegetation shift from dominantly wet evergreen to semi-evergreen at the fossil locality, coincident with an increase in length of the dry season. The comparison of reconstructed CA data and climate modelling data of a Gelasian time slice with that of previously reconstructed climate data by using Climate Leaf Analysis Multivariate Programme (CLAMP) analysis of the late Pliocene–early Pleistocene (Upper Siwalik) fossils of the same locality provides nearly the identical result. Furthermore, all the reconstructed data indicate a monsoonal type of climate during the deposition of the Upper Siwalik sediments.