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Showing papers by "Canadian Forest Service published in 2011"


Journal ArticleDOI
TL;DR: It is shown for the first time successful MPB attack in natural jack pine stands at the leading edge of the epidemic, and a panel of microsatellite loci optimized for both species to classify lodgepole pine, jack pine and their hybrids using simulated data is tested.
Abstract: The current epidemic of the mountain pine beetle (MPB), an indigenous pest of western North American pine, has resulted in significant losses of lodgepole pine. The leading edge has reached Alberta where forest composition shifts from lodgepole to jack pine through a hybrid zone. The susceptibility of jack pine to MPB is a major concern, but there has been no evidence of host-range expansion, in part due to the difficulty in distinguishing the parentals and their hybrids. We tested the utility of a panel of microsatellite loci optimized for both species to classify lodgepole pine, jack pine and their hybrids using simulated data. We were able to accurately classify simulated individuals, and hence applied these markers to identify the ancestry of attacked trees. Here we show for the first time successful MPB attack in natural jack pine stands at the leading edge of the epidemic. This once unsuitable habitat is now a novel environment for MPB to exploit, a potential risk which could be exacerbated by further climate change. The consequences of host-range expansion for the vast boreal ecosystem could be significant.

300 citations


Journal ArticleDOI
TL;DR: Host defenses were major constraints when mountain pine beetle populations were low, but inconsequential after stand-level densities surpassed a critical threshold, while beetles exploited trees weakened by lower-stem insects when populations were too low for cooperative attack.
Abstract: We evaluated the ability of constitutive and inducible defenses to protect trees and restrict herbivore reproduction across the endemic, incipient (i.e., transitory), and eruptive phases of a nativ...

284 citations


Journal ArticleDOI
TL;DR: In this article, the authors summarized recent (2000.2008) rates of extraction, including both conversion and harvest, derived from national forest inventories for North America (the United States, Canada, and Mexico).
Abstract: Incorporating ecological disturbance into biogeochemical models is critical for estimating current and future carbon stocks and fluxes. In particular, anthropogenic disturbances, such as forest conversion and wood harvest, strongly affect forest carbon dynamics within North America. This paper summarizes recent (2000.2008) rates of extraction, including both conversion and harvest, derived from national forest inventories for North America (the United States, Canada, and Mexico). During the 2000s, 6.1 million ha/yr were affected by harvest, another 1.0 million ha/yr were converted to other land uses through gross deforestation, and 0.4 million ha/yr were degraded. Thus about 1.0% of North America fs forests experienced some form of anthropogenic disturbance each year. However, due to harvest recovery, afforestation, and reforestation, the total forest area on the continent has been roughly stable during the decade. On average, about 110 m3 of roundwood volume was extracted per hectare harvested across the continent. Patterns of extraction vary among the three countries, with U.S. and Canadian activity dominated by partial and clear ]cut harvest, respectively, and activity in Mexico dominated by conversion (deforestation) for agriculture. Temporal trends in harvest and clearing may be affected by economic variables, technology, and forest policy decisions. While overall rates of extraction appear fairly stable in all three countries since the 1980s, harvest within the United States has shifted toward the southern United States and away from the Pacific Northwest.

201 citations


Journal ArticleDOI
TL;DR: This approach addresses the data limitation of TLiDAR scans and aims to extract forest architectural metrics at different structural levels.
Abstract: Terrestrial lidar (TLiDAR) has been used increasingly over recent years to assess tree architecture and to extract metrics of forest canopies. Analysis of TLiDAR data remains a difficult task mainly due to the effects of object occlusion and wind on the quality of the retrieved results. We propose to link TLiDAR and tree structure attributes by means of an architectural model. The proposed methodology uses TLiDAR scans combined with allometric relationships to define the total amount of foliage in the crown and to build the tree branching structure. It uses the range (distance) and intensity information of the TLiDAR scans (i) to extract the stem and main branches of the tree, (ii) to reconstruct the fine branching structure at locations where the presence of foliage is very likely, and (iii) to use the availability of light as a criterion to add foliage in the center of the crown where TLiDAR information is sparse or absent due to occlusion effects. An optimization algorithm guides the model towards a realistic tree structure that fits the information gathered from TLiDAR scans and field inventory. The robustness and validity of the proposed model is assessed on five trees belonging to four different conifer species from natural forest environments. This approach addresses the data limitation of TLiDAR scans and aims to extract forest architectural metrics at different structural levels.

190 citations


Journal ArticleDOI
TL;DR: In this paper, the authors experimentally altered soil moisture profiles of peat monoliths collected from several vegetation types common in boreal bogs and used laboratory burn tests to examine the effects of depth-dependent variation in bulk density and moisture on depth of fuel consumption.
Abstract: Theborealbiomeischaracterisedbyextensivewildfiresthatfrequentlyburnintothethickorganicsoilsfound in many forests and wetlands. Previous studies investigating surface fuel consumption generally have not accounted for variation in the properties of organic soils or how this affects the severity of fuel consumption. We experimentally altered soil moisture profiles of peat monoliths collected from several vegetation types common in boreal bogs and used laboratory burn tests to examine the effects of depth-dependent variation in bulk density and moisture on depth of fuel consumption. Depth of burning ranged from 1 to 17cm, comparable with observations following natural wildfires. Individually, fuel bulk density and moisture were unreliable predictors of depth of burning. However, they demonstrated a cumulative influence on the thermodynamics of downward combustion propagation. By modifying Van Wagner's surface fuel consumption model to account for stratigraphic changes in fuel conditions, we were able to accurately predict the maximum depth of fuel consumption for most of the laboratory burn tests. This modified model for predicting the depth of surface fuel consumption in boreal ecosystems may provide a useful framework for informing wildland fire management activities and guiding future development of operational fire behaviour and carbon emission models. Additionalkeywords: bog, boreal,carbon,fire,ground-layerfuels,peat,peatland, smouldering,Sphagnum,surfacefuel combustion.

155 citations


Journal ArticleDOI
TL;DR: A comparison of methods to estimate the loss of carbon from the terrestrial biosphere to the atmosphere from wildland fires is presented in this paper, where the authors identify key uncertainties and areas of improvement for understanding the magnitude and spatial-temporal patterns of pyrogenic carbon emissions across North America.
Abstract: Research activities focused on estimating the direct emissions of carbon from wildland fires across North America are reviewed as part of the North American Carbon Program disturbance synthesis. A comparison of methods to estimate the loss of carbon from the terrestrial biosphere to the atmosphere from wildland fires is presented. Published studies on emissions from recent and historic time periods and five specific cases are summarized, and new emissions estimates are made using contemporary methods for a set of specific fire events. Results from as many as six terrestrial models are compared. We find that methods generally produce similar results within each case, but estimates vary based on site location, vegetation (fuel) type, and fire weather. Area normalized emissions range from 0.23 kg C m−2 for shrubland sites in southern California/NW Mexico to as high as 6.0 kg C m−2 in northern conifer forests. Total emissions range from 0.23 to 1.6 Tg C for a set of 2003 fires in chaparral-dominated landscapes of California to 3.9 to 6.2 Tg C in the dense conifer forests of western Oregon. While the results from models do not always agree, variations can be attributed to differences in model assumptions and methods, including the treatment of canopy consumption and methods to account for changes in fuel moisture, one of the main drivers of variability in fire emissions. From our review and synthesis, we identify key uncertainties and areas of improvement for understanding the magnitude and spatial-temporal patterns of pyrogenic carbon emissions across North America.

141 citations


Journal ArticleDOI
TL;DR: The authors measured changes in carbon (C), nitrogen (N) and phosphorus (P) concentrations and mass of 10 foliar litters decomposing over 12 years at 21 sites across Canada, ranging from subarctic to temperate, to evaluate the influence of litter quality (nature) and forest floor (nurture) on N and P dynamics.
Abstract: We measured changes in carbon (C), nitrogen (N) and phosphorus (P) concentrations and mass of 10 foliar litters decomposing over 12 years at 21 sites across Canada, ranging from subarctic to temperate, to evaluate the influence of litter quality (nature) and forest floor (nurture) on N and P dynamics. Most litters lost P faster than N, relative to C, except in one litter which had a high initial C:P quotient (2,122). Net N loss occurred at mass C:N quotients of between 33 and 68, positively correlated with the C:N quotient in the original litter, and net P loss likely occurred at C:P quotients between 800 and 1,200. Forest floor properties also influenced N and P dynamics: the higher the C:N or C:P quotient in the surface soil organic matter, the smaller the proportion of initial N or P left in the decomposing litter, relative to C. There was a convergence of C:N and C:P quotients as the litters decomposed, with an overall mass ratio of 427:17:1 when the litters reached 20% original C remaining. These results, covering a wide range of sites and litters and thus decomposition rates, showed that the C:N:P quotients followed similar trajectories and converged as the litters decomposed. The relative loss of N and P was affected by both the initial litter nutrient concentration and the chemistry of the site forest floor, with the former being more important than the latter, resulting in spatial variations in nutrient content of the forest floor.

123 citations


Journal ArticleDOI
TL;DR: The cold tolerance strategy and mechanisms involved in the cold tolerance of the emerald ash borer were investigated, and seasonal changes in these mechanisms monitored.

123 citations


Journal ArticleDOI
TL;DR: The authors used difference maps to directly compare downscaled projections of temperature and precipitation across North America for two versions (or generations) of three different Atmospheric-Ocean General Circulation Models (AOGCM)s.
Abstract: Global climate models are constantly being upgraded, but it is often not clear what these changes have on climate change impact projections. We used difference maps to directly compare downscaled projections of temperature and precipitation across North America for two versions (or generations) of three different Atmospheric-Ocean General Circulation Models (AOGCM)s. We found that AOGCM versions differed in their projections for the end of the current century by up to 41C for annual mean temperature and 60% for annual precipitation. To place these changes in an ecological context, we reanalyzed our work on shifts in tree climate envelopes (CEs) using the newer-generation AOGCM projections. Based on the updated AOGCMs, by the 2071–2100 period, tree CEs shifted up to 2.4 degrees further north or 2.6 degrees further south (depending on the AOGCM) and were about 10% larger in size. Despite considerable differences between versions of a given AOGCM, projections made by the newer version of each AOGCM were in general agreement, suggesting convergence across the three models studied here. Assessing the AOGCM outputs in this way provides insight into the magnitude and importance of change associated with AOGCM upgrades as they continue to evolve through time.

121 citations


Journal ArticleDOI
TL;DR: In this paper, the changes in net ecosystem productivity and ecosystem C storage 2-4 and 9-11 years after converting an agricultural land (planted to canola, Brassica napus L.) to hybrid poplar plantations in the Parkland region in central Alberta, Canada.

112 citations


Journal ArticleDOI
TL;DR: In this paper, the authors show that the difference between reflectance measurements in bands associated with the vegetation xanthophyll cycle and estimates of canopy shading obtained from multiangular satellite observations (using the CHRIS/PROBA sensor) permits them to infer plant photosynthetic efficiency, independently of vegetation type and structure (r2 = 0.68, compared to flux measurements).
Abstract: [1] Terrestrial ecosystems absorb about 2.8 Gt C yr?1, which is estimated to be about a quarter of the carbon emitted from fossil fuel combustion. However, the uncertainties of this sink are large, on the order of ±40%, with spatial and temporal variations largely unknown. One of the largest factors contributing to the uncertainty is photosynthesis, the process by which plants absorb carbon from the atmosphere. Currently, photosynthesis, or gross ecosystem productivity (GEP), can only be inferred from flux towers by measuring the exchange of CO2 in the surrounding air column. Consequently, carbon models suffer from a lack of spatial coverage of accurate GEP observations. Here, we show that photosynthetic light use efficiency (?), hence photosynthesis, can be directly inferred from spaceborne measurements of reflectance. We demonstrate that the differential between reflectance measurements in bands associated with the vegetation xanthophyll cycle and estimates of canopy shading obtained from multiangular satellite observations (using the CHRIS/PROBA sensor) permits us to infer plant photosynthetic efficiency, independently of vegetation type and structure (r2 = 0.68, compared to flux measurements). This is a significant advance over previous approaches seeking to model global-scale photosynthesis indirectly from a combination of growth limiting factors, most notably pressure deficit and temperature. When combined with modeled global-scale photosynthesis, satellite-inferred ? can improve model estimates through data assimilation. We anticipate that our findings will guide the development of new spaceborne approaches to observe vegetation carbon uptake and improve current predictions of global CO2 budgets and future climate scenarios by providing regularly timed calibration points for modeling plant photosynthesis consistently at a global scale.

Journal ArticleDOI
TL;DR: In this article, the authors used wavelet analysis to synthesize the performance of 21 ecosystem models at 9 eddy covariance towers as part of the North American Carbon Program's site-level intercomparison.
Abstract: [1] Ecosystem models are important tools for diagnosing the carbon cycle and projecting its behavior across space and time. Despite the fact that ecosystems respond to drivers at multiple time scales, most assessments of model performance do not discriminate different time scales. Spectral methods, such as wavelet analyses, present an alternative approach that enables the identification of the dominant time scales contributing to model performance in the frequency domain. In this study we used wavelet analyses to synthesize the performance of 21 ecosystem models at 9 eddy covariance towers as part of the North American Carbon Program's site-level intercomparison. This study expands upon previous single-site and single-model analyses to determine what patterns of model error are consistent across a diverse range of models and sites. To assess the significance of model error at different time scales, a novel Monte Carlo approach was developed to incorporate flux observation error. Failing to account for observation error leads to a misidentification of the time scales that dominate model error. These analyses show that model error (1) is largest at the annual and 20–120 day scales, (2) has a clear peak at the diurnal scale, and (3) shows large variability among models in the 2–20 day scales. Errors at the annual scale were consistent across time, diurnal errors were predominantly during the growing season, and intermediate-scale errors were largely event driven. Breaking spectra into discrete temporal bands revealed a significant model-by-band effect but also a nonsignificant model-by-site effect, which together suggest that individual models show consistency in their error patterns. Differences among models were related to model time step, soil hydrology, and the representation of photosynthesis and phenology but not the soil carbon or nitrogen cycles. These factors had the greatest impact on diurnal errors, were less important at annual scales, and had the least impact at intermediate time scales.

Journal ArticleDOI
TL;DR: This article reviewed the historical context and the research conducted over the past 15 years that has led to the province's current vegetation management strategy and identified the major challenges of vegetation management being faced in Quebec in the context of intensive silviculture and ecosystem-based management.
Abstract: Vegetation management is crucial to meeting the objectives of forest plantations. Following public hearing processes, chemical herbicides were banned on Crown forest lands in Quebec (Canada) in 2001. Release now mainly relies on mechanical treatments. Our objectives are to review the historical context and the research conducted over the past 15 years that has led to the province’s current vegetation management strategy and to identify the major challenges of vegetation management being faced in Quebec in the context of intensive silviculture and ecosystem-based management. Research has led to an integrated management model without herbicides, adapted to the ecological characteristics of reforestation sites. The Quebec experience illustrates how, on most sites, vegetation management that is based on early reforestation, the use of tall planting stock and intensive mechanical release brings crop trees to the free-to-grow stage without the use of herbicides and without resulting in major effects on vegetation diversity. This vegetation management strategy is an asset in the implementation of ecosystem-based management. However, research demonstrates that mechanical release alone does not promote optimal crop-tree growth, due to rapid resprouting or suckering of competitors and competition from herbaceous species. Therefore, the current strategy poses important challenges in the management of plantations where the objective is to maximise wood production.

Journal ArticleDOI
TL;DR: A decadal-scale multiproxy record of minerals, pollen, and charcoal from Kettle Lake, North Dakota provides a high-resolution record of climate and vegetation change spanning the entire Holocene from the northern Great Plains (NGP) in North America.

Journal ArticleDOI
TL;DR: In this paper, two parametric approaches, the bootstrap and the jackknife, were compared to a parametric estimator for estimating uncertainty using the k-NN technique with forest inventory and Landsat data from Finland, Italy, and the USA.

Journal ArticleDOI
TL;DR: This article provided the first national assessment of wildfire-related evacuations in Canada and documents the loss of homes that coincided with evacuation events, finding that despite the intensity and abundance of wildfire in Canada, wildfires have displaced a relatively small number of people.
Abstract: Evacuations represent an integral aspect of protecting public safety in locations where intense, fast-spreading forest fires co-occur with human populations. Most Canadian fire management agencies have as their primary objective the protection of people and property, and all fire management agencies in Canada recommend evacuations when public safety is in question. This study provides the first national assessment of wildfire-related evacuations in Canada and documents the loss of homes that coincided with evacuation events. The most striking finding is that despite the intensity and abundance of wildfire in Canada, wildfires have displaced a relatively small number of people. Between 1980 and 2007, the median number of evacuees and home losses per year in Canada were 3,590 and 2, respectively. Evacuees’ homes survived in 99.3% of cases. Patterns of evacuations and home losses reflected the distributions of forests, wildfire, and people across the Canadian landscape. Most evacuations occurred in boreal areas, which have relatively low population densities but among the highest percent annual area burned in Canada. Evacuations were less common in southern parts of the country, where most Canadians reside, but individual wildfires in these areas had significant impacts. Interactions between wildfire and people in Canada exhibited a unique regional pattern, and within the most densely populated regions of the country they can be considered ‘low-probability, high-consequence’ events. This Canadian context is fundamentally different from places such as California, where concentrations of fires and people overlap across large areas and therefore calls for a fundamentally different fire management response.

Journal ArticleDOI
TL;DR: Research indicates that branch sampling greatly reduces false negatives associated with visual surveys and window sampling at breast height, and could be incorporated into routine sanitation or maintenance of city-owned trees to identify and delineate infested areas.
Abstract: The emerald ash borer, Agrilus planipennis Fairmaire, is an exotic invasive insect causing extensive mortality to ash trees, Fraxinus spp, in Canada and the United States Detection of incipient populations of this pest is difficult because of its cryptic life stages and a multiyear time lag between initial attack and the appearance of signs or symptoms of infestation We sampled branches from open-grown urban ash trees to develop a sample unit suitable for detecting low density A planipennis infestation before any signs or symptoms are evident The sample unit that maximized detection rates consisted of one 50-cm-long piece from the base of a branch ≥6 cm diameter in the midcrown The optimal sample size was two such branches per tree This sampling method detected ≈75% of asymptomatic trees known to be infested by using more intensive sampling and ≈3 times more trees than sampling one-fourth of the circumference of the trunk at breast height The method is less conspicuous and esthetically damaging to a tree than the removal of bark from the main stem or the use of trap trees, and could be incorporated into routine sanitation or maintenance of city-owned trees to identify and delineate infested areas This research indicates that branch sampling greatly reduces false negatives associated with visual surveys and window sampling at breast height Detection of A planipennis-infested asymptomatic trees through branch sampling in urban centers would provide landowners and urban foresters with more time to develop and implement management tactics

Journal ArticleDOI
TL;DR: Results from this study highlight the importance of field research and the need to include multiple endpoints when examining potential effects of a contaminant on non-target organisms.

Journal ArticleDOI
TL;DR: In this article, the ecosys model was applied to investigate the effects of water table and subsurface hydrology changes on carbon dioxide exchange at the ombrotrophic Mer Bleue peatland, Ontario, Canada.
Abstract: [1] The ecosys model was applied to investigate the effects of water table and subsurface hydrology changes on carbon dioxide exchange at the ombrotrophic Mer Bleue peatland, Ontario, Canada. It was hypothesized that (1) water table drawdown would not affect vascular canopy water potential, hence vascular productivity, because roots would penetrate deeper to compensate for near‐surface dryness, (2) moss canopy water potential and productivity would be severely reduced because rhizoids occupy the uppermost peat that is subject to desiccation with water table decline, and (3) given that in a previous study of Mer Bleue, ecosystem respiration showed little sensitivity to water table drawdown, gross primary productivity would mainly determine the net ecosystem productivity through these vegetation–subsurface hydrology linkages. Model output was compared with literature reports and hourly eddy‐covariance measurements during 2000–2004. Our findings suggest that late‐summer water table drawdown in 2001 had only a minor impact on vascular canopy water potential but greatly impacted hummock moss water potential, where midday values declined to −250 MPa on average in the model. As a result, simulated moss productivity was reduced by half, which largely explained a reduction of 2–3 mmol CO2 m −2 s −1 in midday simulated and measurement‐derived gross primary productivity and an equivalent reduction in simulated and measured net ecosystem productivity. The water content of the near‐surface peat (top 5–10 cm) was found to be the most important driver of interannual variability of annual net ecosystem productivity through its effects on hummock moss productivity and on ecosystem respiration.

Journal ArticleDOI
TL;DR: The results are used to parameterize a maximum likelihood estimation model of temperature-dependent instar IV developmental rates, including the effect of diapause, and can be included as part of a spruce beetle phenology model for predicting population dynamics.

Journal ArticleDOI
TL;DR: In this paper, a 3-parameter model (DOC-3) was proposed to project daily stream DOC concentrations and fluxes from modelled estimates for daily soil temperature and moisture, year-round, and in relation to basin size and wetness.

Journal ArticleDOI
TL;DR: In this article, a significant increase in the warming trend observed in recent decades in semi-arid Mongolia, tree-ring attributes and anomalies were analyzed to detect potential changes in the growth-climate relationship.
Abstract: In light of a significant increase in the warming trend observed in recent decades in semi-arid Mongolia, tree-ring attributes and anomalies were analysed to detect potential changes in the growth-climate relationship. In a moisture-limited environment, an increase in temperature could cause a shift in the seasonal response of trees to climate. Chronologies were developed for the dominant tree species (Larix sibirica Ledeb., Pinus sibirica Du Tour, and Pinus sylvestris L.) from north- central Mongolia. In addition to annual ring width, both earlywood and latewood width were measured, and tree-ring anomalies such as false rings and light rings were systematically identified. Earlywood width was mainly associated with precipitation in the year prior to ring formation and early growing season conditions. Temperature was associated with cur- rent year growth and mainly influenced latewood development. False rings were good indicators of early summer droughts, whereas light rings were mainly associated with a cold end of summer. A seasonal shift in the significance of monthly cli- mate variables was observed in recent decades. This displacement presumably resulted from changes in the timing and dura- tion of the growing season. Tree growth starts earlier in spring and is now affected by late summer to early autumn climate conditions. Resume : En lien avec la tendance significative au rechauffement observee depuis les dernieres decennies en Mongolie semi-aride, des attributs et des anomalies des cernes annuels ont ete analyses pour detecter les changements potentiels dans la relation entre la croissance des arbres et le climat. Dans des milieux ou la disponibilite en eau est un facteur limitant, des augmentations de temperature pourraient engendrer, une modification de la reponse saisonniere des arbres au climat. Des se- ries dendrochronologiques ont ete developpees pour les trois especes dominantes du centre-nord de la Mongolie (Larix sibi- rica Ledeb., Pinus sibirica Du Tour et Pinus sylvestris L.). En plus de la largeur des cernes annuels, la largeur du bois initial et du bois final a ete mesuree et des anomalies des cernes telles que les faux-cernes et les cernes pâles ont ete syste- matiquement identifiees. La largeur du bois initial etait principalement associee aux precipitations de la saison precedant la formation du cerne ainsi qu'a celles du debut de la saison de croissance. La saison de croissance en cours etait pour sa part associee a la temperature qui influencait surtout le developpement du bois final. Les faux cernes etaient de bons indicateurs de secheresses en debut d'ete, tandis que les cernes pâles etaient surtout associes a des fins d'ete froides. Une modification saisonniere des associations significatives avec les variables climatiques mensuelles a ete observee au cours des dernieres decennies. Cette modification pourrait etre reliee a des changements dans l'initiation et la duree de la saison de croissance. La croissance des arbres commence plus tot au printemps et est maintenant influencee par les conditions climatiques qui prevalent a la fin de l'ete et au debut de l'automne.

Journal ArticleDOI
TL;DR: In this article, the authors used the process model ecosys to test hypotheses for these changes with CO2 and energy fluxes measured by eddy covariance over a mesic shrub tundra at Daring Lake, Canada, under varying growing seasons.
Abstract: [1] Changes in arctic C stocks with climate are thought to be caused by rising net primary productivity (NPP) during longer and warmer growing seasons, offset by rising heterotrophic respiration (Rh) in warmer and deeper soil active layers. In this study, we used the process model ecosys to test hypotheses for these changes with CO2 and energy fluxes measured by eddy covariance over a mesic shrub tundra at Daring Lake, Canada, under varying growing seasons. These tests corroborated substantial rises in NPP, smaller rises in Rh, and, hence, rises in net ecosystem productivity (NEP) from 17 to 45 g C m−2 yr−1 (net C sink), modeled with higher Ta and longer growing seasons. However, NEP was found to decline briefly during midsummer warming events (Ta > 20°C). A model run under climate change predicted for Daring Lake indicated that rises in NPP would exceed those in Rh during the first 100 years, causing NEP to rise. Rises in NPP were driven by more rapid net N mineralization from more rapid Rh in warming soils. However, greater declines in NEP were modeled during more frequent and intense midsummer warming events as climate change progressed. Consequently, average annual NEP (± interannual variability) rose from 30 (±13) g C m−2 yr−1 under current climate to 57 (±40) g C m−2 yr−1 after 90 years but declined to 44 (±51) g C m−2 yr−1 after 150 years, indicating that gains in tundra NEP under climate change may not be indefinite.

Journal ArticleDOI
TL;DR: It is suggested that Z3-6:OH—baited green traps placed in the ash canopy would be a superior lure for detecting and monitoring A. planipennis throughout the flight season.
Abstract: We conducted trapping experiments for the emerald ash borer, Agrilus planipennis Fairmaire (Coleoptera: Buprestidae) in Michigan, USA, and Ontario, Canada, to compare unbaited light green sticky prism traps with traps baited with phoebe oil, (Z)-3-hexenol (Z3-6:OH), or blends of other green leaf volatiles (GLVs) with Z3-6:OH. Traps were placed in the lower canopy of ash trees (Fraxinus spp.). Catches with Z3-6:OH—baited traps showed a significant male bias and these traps caught significantly more males than the unbaited controls at both sites. They were also superior to phoebe oil-baited traps and those baited with GLV blends. Catches with phoebe oil showed a significant female bias but there was no difference in the number of females captured between traps baited with phoebe oil or Z3-6:OH lures. Catches were analyzed at regular time intervals to examine the response of A. planipennis to the lures over the course of the flight season. Z3-6:OH—baited traps consistently caught more males than the...

Journal ArticleDOI
TL;DR: There was no significant difference in growth rate among NA1 wild type (wt), NA2, and EU1 lineages at any temperature tested and the NA1 lineage of P. ramorum was the most variable in aggressiveness and growth rate.
Abstract: Summary There are three major clonal lineages of Phytophthora ramorum present in North America and Europe named NA1, NA2, and EU1. Twenty-three isolates representing all three lineages were evaluated for phenotype including (i) aggressiveness on detached Rhododendron leaves and (ii) growth rate at minimum, optimum, and maximum temperatures. Closely related species P. foliorum and P. hibernalis were included in phenotypic tests since these species are encountered in nursery surveys for P. ramorum. Isolates from the NA2 and EU1 lineages were the most aggressive and isolates from the NA1 group were the least aggressive. The NA1 lineage of P. ramorum was the most variable in aggressiveness and growth rate. The variability in the NA1 lineage was due to the presence of non-wild type (nwt) isolates. There was no significant difference in growth rate among NA1 wild type (wt), NA2, and EU1 lineages at any temperature tested. The difference between wt and nwt P. ramorum isolates is discussed.

Journal ArticleDOI
01 Jan 2011-Forestry
TL;DR: Although the Big Qualicum (East Vancouver Island) population had the same high resistance as the Haney (Fraser Valley) population, it was expressed primarily through increased CRC, and sclereid cell density had the strongest correlation to weevil attack followed by CRC.
Abstract: Summary It has long been known that strong expressions of resistance to the white pine weevil ( Pissodes strobi Peck) exist in certain Sitka spruce ( Picea sitchensis (Bong.) Carr.) populations, particularly among trees originating from the Fraser Valley and the Qualicum area of British Columbia (BC). In this paper , we characterize how resistance is manifested in these known resistant populations. Specifi cally, using cloned individuals, we investigated resistant traits associated with repellency , constitutive resin canals (CRC) and sclereid or stone cells. Results indicate signifi cant population differences in the level of these traits between these two populations and susceptible populations. Fraser Valley populations had four times the sclereid density of susceptible populations. Although the Big Qualicum (East Vancouver Island) population had the same high resistance as the Haney (Fraser Valley) population, it was expressed primarily through increased CRC. Sclereid cell density had the strongest correlation to weevil attack followed by CRC. We discuss pathways by which two distinct resistant populations may have developed in this high weevil hazard region of south - west BC.

Journal ArticleDOI
NaultJ.1
TL;DR: Water-extracted soil concentrations of NH4+ were up to 1.5 times greater than NO3– averaged across sites, and the amino acid N to inorganic N ratio was up to 2.4:1 in soils under Vaccinium.
Abstract: There are few examinations of the relative availability and plant uptake of inorganic N and amino acid N in temperate forest regions. We determined the availability of amino acid N and inorganic N ...

Journal ArticleDOI
TL;DR: Logging is the main human disturbance in the boreal forest; thus, understanding the effects of harvesting practices on biodiversity is essential for a more sustainable forestry and further efforts toward the understanding of species composition in higher strata of the Boreal forest are needed.
Abstract: Logging is the main human disturbance in the boreal forest; thus, understanding the effects of harvesting practices on biodiversity is essential for a more sustainable forestry. To assess changes in spider composition because of harvesting, samples were collected from three forest layers (overstory, understory, and ground) of deciduous and conifer dominated stands in the northwestern Canadian boreal mixedwood forest. Spider assemblages and feeding guild composition were compared between uncut controls and stands harvested to 20% retention. In total, 143 spider species were collected, 74 from the ground, 60 from the understory, and 71 from the overstory, and species composition of these three pools differed considerably among layers. Distinctive spider assemblages were collected from the canopy of each forest cover type but these were only slightly affected by harvesting. However, logging had a greater impact on the species composition in the understory and ground layers when compared with unharvested controls. Guild structure differed among layers, with wandering and sheet-weaving spiders dominant on the ground while orb-weaving and ambush spiders were better represented in the understory and overstory, respectively. Given the ecological importance of spiders and the expectation of faunal changes with increased harvesting, further efforts toward the understanding of species composition in higher strata of the boreal forest are needed.

Journal ArticleDOI
TL;DR: An efficient method for assessing lichen biomass at the stand level in boreal forests is presented and it accounts for the variation in biomass among lichen species that are equally abundant.
Abstract: Lichens are an important winter food source for woodland caribou (Rangifer tarandus caribou), but quantifying their abundance is difficult. Here, we present an efficient method for assessing lichen biomass at the stand level in boreal forests. We measured lichens occurring in high enough abundance to serve as a winter food source for woodland caribou in 51 boreal forest stands. Samples of each species or genus were collected from each stand and a mean abundance (cover) to biomass ratio was established. The method does not require samples to be collected or weighed, due to this predetermined relationship, and it also accounts for the variation in biomass among lichen species that are equally abundant. The variation in lichen growth between stands was assessed by means of five lichen abundance classes. The proposed method was tested in 34 stands with a wide range of ages and stem densities. The average time to complete a lichen biomass assessment was approximately 2 h. This method is an efficient and accura...

Journal ArticleDOI
TL;DR: In this article, the association between hemispheric-scale climate patterns and annual wildfire activity can be obscured by l o nth-order climate change, which is regulated in part by climate.
Abstract: Wildfire impacts on ecological and socioeconomic systems are regulated, in part, by climate. Association between hemispheric-scale climate patterns and annual wildfire activity can be obscured by l...